Paleobiota of the Green River Formation explained

The Green River Formation is a geological formation located in the Intermountain West of the United States, in the states of Colorado, Utah, and Wyoming. It comprises sediments deposited during the Early Eocene in a series of large freshwater lakes: Lake Gosiute, Lake Uinta, and Fossil Lake (the last containing Fossil Butte National Monument). It preserves a high diversity of freshwater fish, birds, reptiles, and mammals, with some sections of the formation (including Fossil Lake and the Parachute Creek member of Lake Uinta) qualifying as Konservat-Lagerstätten due to their extremely well-preserved fossils.

Cartilaginous fish

Family	Genus	Species	Member	Abundance	Notes	Image
†Asterotrygonidae	†Asterotrygon	†A. maloneyi	Fossil Lake		An Asterotrygonidae stingray of uncertain higher affinities.[1] [2]
†Heliobatidae	†Heliobatis	†H. radians	Fossil Lake		A Heliobatid stingray of uncertain higher affinities.

Bony fish

Primarily based on Grande (2001), with changes where necessary:

Lepisosteiformes

Family	Genus	Species	Member	Abundance	Notes	Image
Lepisosteidae	Atractosteus	†A. atrox	Fossil Lake		An alligator gar relative. A. atrox is the largest Green River gar.
		†A. simplex	Fossil Lake Lake Gosiute?		An alligator gar relative.
	†Cuneatus	†C. cuneatus	Fossil Lake? Lake Gosiute Lake Uinta		A Cuneatini tribe gar.
		†C. wileyi	Lake Uinta		A Cuneatini tribe gar.
	Lepisosteus	†L. bemisi	Fossil Lake		A Lepisosteini tribe gar.[3]
	†Masillosteus	†M. janeae	Fossil Lake		A Cuneatini tribe gar.

Amiiformes

Family	Genus	Species	Member	Abundance	Notes	Image
Amiidae	Amia	†A. pattersoni	Fossil Lake Lake Gosiute		An amiid bowfin.
	†Cyclurus	†C. gurleyi	Fossil Lake	Only 8 specimens known	An amiid related to modern bowfins.

Osteoglossiformes

Family	Genus	Species	Member	Abundance	Notes	Image
Osteoglossidae	†Phareodus	†P. encaustus	Fossil Lake Lake Gosiute		A phareodontine bonytongue.
		†P. testis	Fossil Lake

Clupeiformes

Family	Genus	Species	Member	Abundance	Notes	Image
Clupeidae	†Gosiutichthys	†G. parvus	Lake Gosiute		A clupeine clupeid herring relative.[4]
	†Knightia	†K. alta	Fossil Lake Lake Gosiute Lake Uinta		A pellonuline clupeid herring relative.[5]
		†K. eocaena	Fossil Lake Lake Gosiute Lake Uinta	The most common articulated vertebrate fossil worldwide.[6]	A pellonuline clupeid herring relative.

Siluriformes

Family	Genus	Species	Member	Abundance	Notes	Image
†Hypsidoridae	†Hypsidoris	†H. farsonensis	Lake Gosiute		A hypsidorid catfish.
Ictaluridae	†Astephus	†A. antiquus	Fossil Lake Lake Gosiute Lake Uinta		An ictalurid catfish.

Percopsiformes

Family	Genus	Species	Member	Abundance	Notes	Image
Percopsidae	†Amphiplaga	†A. brachyptera	Fossil Lake		A trout-perch relative. Includes the jr synonym Erismatopterus endlichi (1877)
	†Erismatopterus	†E. levatus	Lake Gosiute Lake Uinta		A trout-perch relative. First described as Cyprinodon levatus (1870) includes the jr synonym Erismatopterus rickseckeri (1871)

Acanthuriformes

Family	Genus	Species	Member	Abundance	Notes	Image
Moronidae	†Cockerellites	†C. liops	Fossil Lake	common	A planktivorous/insectivorous temperate bass. First described as Priscacara liops (1877) First moved to Cockerellites liops in 1923.
	†Hypsiprisca	†H. hypsacantha	Fossil Lake	rare	A temperate bass. First named Priscacara hypsacantha 1886)
		†H. sp.	Fossil Lake	rare	An undescribed species of Hypsiprisca. A slenderer species with a more rounded tail fin outline.
	†Priscacara	†P. serrata	Fossil Lake Lake Uinta		A temperate bass.
		†P. sp. a	Fossil Lake	1 specimen	An undescribed Priscacara Specimens have a rounder body and weak dorsal spines.
		†P. sp. b.	Fossil Lake	1 specimen	An undescribed Priscacara Possibly a new species or a variant specimen with 4 anal spines.

Incertae sedis

Family	Genus	Species	Member	Abundance	Notes	Image
†Asineopidae	†Asineops	†A. squamifrons	Fossil Lake Lake Gosiute Lake Uinta		An acanthomorph of uncertain affinities Possibly a percopsiform, polymixiiform, or perciform.
		A. sp.	Fossil Lake		An undescribed Asineops species Restricted to Fossil Lake.

Amphibians

Salamanders

Reptiles

Squamates

Family	Genus	Species	Member	Abundance	Notes	Image
Boidae	†Boavus	†B. idelmani	Fossil Lake		A boa.
Dactyloidae	†Afairiguana	†A. avius	Fossil Lake		A stem-anole.[10]
Shinisauridae	†Bahndwivici	†B. ammoskius	Fossil Lake		A relative of the Chinese crocodile lizard.
Tropidophiidae?	†Dunnophis	D. sp.	Fossil Lake		An undescribed possible dwarf boa.[11]
Varanidae	†Saniwa	†S. ensidens	Fossil Lake		A stem-monitor lizard.
Incertae sedis	Indeterminate	Indeterminate	Lake Uinta (Parachute Creek)	1 specimen	A possible gecko or iguanid Preserved with soft tissue and skin, but bones dissolved.[12]
Incertae sedis	Indeterminate	Indeterminate	Lake Uinta (Parachute Creek)	1 specimen	A relative of Shinisaurids Preserved with soft tissue and skin, but bones dissolved.[13]

Crocodilians

Family	Genus	Species	Member	Abundance	Notes	Image
Alligatoridae	†Tsoabichi	†T. greenriverensis	Fossil Lake		A caiman.[14]
Incertae sedis	†Borealosuchus	†B. wilsoni	Fossil Lake		A eusuchian crocodyliform of uncertain affinities. Moved from Leidyosuchus wilsoni (1959)
		Cf. †B. sp.	Fossil Lake		Indeterminate eusuchian bones close to Borealosuchus.
	"Crocodylus"		Lake Uinta (Manti beds)	1 specimen	A Crocodyloidea of unplaced affinity. Not a Crocodylus species, but not given a new genus as of 2024.[15]
			Lake Gosiute		Bones close in morphology to †"Crocodylus" affinis A Crocodyloidea of unplaced affinity. Not a Crocodylus species, but not given a new genus as of 2024.

Turtles

Family	Genus	Species	Member	Abundance	Notes	Image
Baenidae	†Chisternon	†C. undatum	Fossil Lake		A baenid.
Dermatemydidae	†Baptemys	†B. wyomingensis	Fossil Lake		A dermatemydid.
		†B. Cf. wyomingensis	Lake Unita		A dermatemydid.
Geoemydidae	†Echmatemys	†E. haydeni		1 specimen	A geoemydid tortoise.[16]
		†E. naomi	Fossil Lake	1 specimen	A geoemydid tortoise.[17]
		†E. wyomingensis	Fossil Lake		A geoemydid tortoise.[18]
Testudinidae	†Hadrianus	†Cf. †H. corsoni	Fossil Lake		A tortoise species. A single complete hatchling
Trionychidae	†Axestemys	A. sp.	Fossil Lake		A softshell turtle.
	†Oliveremys	†O. uintaensis	Fossil Lake		A softshell turtle.
	†Plastomenus	†P. thomasii	Fossil Lake		A softshell turtle.

Birds

Lithornithiformes

Family	Genus	Species	Member	Abundance	Notes	Image
†Lithornithidae	†Calciavis	†C. grandei	Fossil Lake		A lithornithid.[19]
	†Pseudocrypturus	†P. cercanaxius	Fossil Lake		A lithornithid.

Anseriformes

Family	Genus	Species	Member	Abundance	Notes	Image
†Presbyornithidae	†Presbyornis	†P. pervetus	Fossil Lake		A presbyornithid anseriform.
Incertae sedis	†Paakniwatavis	†P. grandei	Fossil Lake		An anseriform of uncertain affinities.

Leptosomiformes

Family	Genus	Species	Member	Abundance	Notes	Image
incertae sedis	†Plesiocathartes	P. major	Fossil Lake		A stem-cuckoo roller.[20]
		†P. wyomingensis	Fossil Lake		A stem-cuckoo roller

Coraciiformes

Family	Genus	Species	Member	Abundance	Notes	Image
Primobucconidae	†Primobucco	†P. mcgrewi	Fossil Lake		A primobucconid coraciiform.
Incertae sedis	†Paracoracias	†P. occidentalis			A coraciiform.[21]

Piciformes

Family	Genus	Species	Member	Abundance	Notes	Image
Incertae sedis	†Neanis	†N. schucherti	Fossil Lake		A possible piciform of uncertain affinities.
	†Pristineanis	†P. kistneri			A possible piciform of uncertain affinities.[22]

Strisores

Family	Genus	Species	Member	Abundance	Notes	Image
Eocypselidae	†Eocypselus	†E. rowei	Fossil Lake		A eocypselid apodiform.[23]
Incertae sedis	†Fluvioviridavis	†F. platyrhamphus	Fossil Lake		A basal strisore, possibly a stem-frogmouth.[24] [25]
	†Prefica	†P. nivea	Fossil Lake		A stem-oilbird.[26]

Suliformes

Family	Genus	Species	Member	Abundance	Notes	Image
Fregatidae	†Limnofregata	L. azygosternon	Fossil Lake		An early frigatebird.
		L. hasegawai	Fossil Lake		An early frigatebird

Pelecaniformes

Family	Genus	Species	Member	Abundance	Notes	Image
Threskiornithidae	†Rhynchaeites	†R. sp.		1 specimen	An undescribed ibis, known from a largely complete specimen.
	†Vadaravis	†V. brownae	Fossil Lake		Potentially a stem-ibis, but may be an indeterminate aequornithine.[28]

Eufalconimorphae

Family	Genus	Species	Member	Abundance	Notes	Image
†Halcyornithidae	†Cyrilavis	†C. colburnorum	Fossil Lake		A halcyornithid.
		†C. olsoni	Fossil Lake
†Messelasturidae	†Tynskya	†T. eocaena	Fossil Lake		A messelasturid.
†Psittacopedidae	†Eofringillirostrum	†E. boudreauxi	Fossil Lake		A psittacopedid.[30]
†Quercypsittidae	†Avolatavis	†A. tenens	Fossil Lake		A quercypsittid.
†Zygodactylidae	†Eozygodactylus	†E. americanus	Fossil Lake		A zygodactylid.[31]
	†Zygodactylus	†Z. grandei	Fossil Lake		A zygodactylid.[32]

Neoaves incertae sedis

Two other genera, "Eoeurypyga" (a stem-sunbittern) and "Wyomingcypselus" (an early apodiform) are mentioned only in a 2002 dissertation,^[34] and are presently nomina nuda.

Avian ichnofossils

Family	Genus	Species	Member	Abundance	Notes	Image
†Avipedidae	†Avipeda	Cf. †A. phoenix	Lake Unita		Avian trackways with three forward facing toes without inter-digit webbing.[35]
†Ignotornidae	†Quadridigitus	†Q. semimembranus	Lake Unita		Avian trackways with three forward facing toes, a hallux, and inter-digit webbing.
†Jindongornipodidae	†Jindongornipes	†J. falkbuckleyi	Lake Unita		Large avian trackways with four toes, and wide spread between digits II and IV.
Incertae sedis	†Presbyornithiformipes	†P. feduccii	Lake Uinta		Duck-like trackways with dabble marks attributed to Presbyornis.[36]

Mammals

Partially based on Grande (1984). Aside from the few well-preserved mammals found in Fossil Lake, a majority of Green River mammals are based on isolated bones and teeth:

Metatheria

Family	Genus	Species	Member	Abundance	Notes	Image
†Herpetotheriidae	†Herpetotherium	†H. innominatum	Lake Gosiute Lake Uinta		An opossum-like basal metatherian Originally described as Peratherium innominatum.
		†H. knighti	Lake Gosiute		An opossum-like basal metatherian.

Chiroptera

Family	Genus	Species	Member	Abundance	Notes	Image
†Icaronycteridae	†Icaronycteris	†I. gunnelli	Fossil Lake	2 specimens	An icaronycterid bat.[37]
		†I. index	Fossil Lake	2 specimens	An icaronycterid bat.
†Onychonycteridae	†Onychonycteris	†O. finneyi	Fossil Lake	2 specimens	An onychonycterid bat.

Eulipotyphla

family	Genus	Species	Member	Abundance	Notes	Image
Nyctitheriidae	Nyctitherium	N. nitidum	Lake Gosiute		A nyctitheriid. Originally described as Talpavus nitidus.
		N. sp.	Lake Uinta

Pan-Carnivora

Family	Genus	Species	Member	Abundance	Notes	Image
Miacidae	Vulpavus	V. profectus	Lake Gosiute		A miacid.
	Indeterminate	indeterminate	Lake Gosiute Lake Uinta		Indeterminate miacid bones.[38]
Sinopidae	Sinopa	S. minor	Lake Uinta		A sinopid hyaenodont.
Viverravidae	Viverravus	V. minutus	Lake Uinta		A viverravid.
Incertae sedis	"Miacis"	"M." gracilis	Lake Uinta		A carnivoraform of uncertain affinities.

Pan-Perissodactyla

family	Genus	Species	Member	Abundance	Notes	Image
†Brontotheriidae	†Lambdotherium	†L. popoagicum	Fossil Lake		A small-sized basal brontothere.
	†L. sp.	Lake Unita		A small-sized basal brontothere. Reported from Powder Wash, Utah by Cope (1880)[40]
Equidae	†Protorohippus	†Aff. P. venticolus	Fossil Lake	2 specimens	An equid closely affiliated with P. venticolus.[41]
	Indeterminate	Indeterminate	Lake Gosiute Lake Unita<	Indeterminate equiid bones. First identified as Hyracotherium sp.
†Hyopsodontidae	†Hyopsodus	†H. minusculus	Lake Gosiute Lake Uinta		A hyopsodontid.
		†H. paulus (=H. vicarius)	Lake Uinta		A hyopsodontid. includes H. vicarius specimens.
		†H. wortmani	Fossil Lake		A hyopsodontid.
↑Hyrachyidae	↑Hyrachyus	Indeterminate	Lake Unita		Undeterminate Hyrachyus bones.
Indeterminate	Indeterminate	Indeterminate	Fossil Lake	1 specimen	A juvenile tapiromorph, the largest mammal known from the formation. Initially described as Heptodon calciculus.[42]

Rodentia

Family	Genus	Species	Member	Abundance	Notes	Image
Ischyromyidae	Paramys	P. sp.	Lake Uinta		A ischyromyid rodent.
	Thisbemys	T. sp.	Lake Gosiute		An undescribed ischyromyid rodent.
Sciuravidae	Sciuravus	S. eucristadens	Lake Uinta (Douglas Creek Member)		A sciuravid rodent. Originally described as Viverravus eucristadens (1937).
		S. sp.	Lake Uinta		An undescribed sciuravid.

Primatomorpha

Family	Genus	Species	Member	Abundance	Notes	Image
Microsyopidae	Microsyops	M. elegans	Lake Gosiute		A microsyopid stem-primate.
	Uintasorex	U. parvulus	Lake Gosiute		A microsyopid stem-primate.
Notharctidae	Notharctus	N. sp.	Lake Gosiute		A notharctid adapiform.
	Smilodectes	S. gracilis	Lake Uinta		A notharctid adapiform. Formerly included in Notharctus matthewi.
Omomyidae	Anemorhysis	A. sp.	Lake Unita	1 specimen (lost?)	An omomyid primate. First named as Uintalacus nettingi (1958) Deemed Anemorhysis sp indeterminate (1976)[44]
	Omomys	O. carteri	Lake Uinta		An omomyid primate. Includes O. pucillus[45]
	Tetonius	T. sp.	Lake Uinta		An omomyid primate.
	Washakius	W. insignis	Lake Gosiute		An omomyid primate.

Arthropoda

Arachnida

Araneae

Family	Genus	Species	Member	Abundance	Notes	Image
Araneidae	Araneus	A. sp	Lake Gosiute	6 specimens	Possible orbweb-spiders, known from partial fossils First placed as Epeira sp.[46]
Hersiliidae	Hersiliola (sensu lato)	H. sp.	Lake Uinta (Parachute Creek Member)	1 male	A tree trunk spider.[47]
Selenopidae	Indeterminate	Indeterminate	Lake Uinta (Parachute Creek Member)	1 specimen	A female or juvenile selenopid spider.
Thomisidae	Indeterminate	Indeterminate	Lake Uinta (Parachute Creek Member)	2 males, 1 juv female	Crab spiders.
Uloboridae	Miagrammopes	M. sp.	Lake Uinta (Parachute Creek Member)	1 female, 2 males	A uloborid spider.
	Indeterminate	Indeterminate	Lake Uinta (Parachute Creek Member)		A uloborid spider.
Unidentified	†Araneaovoius	†A. columbiae[48]		1 specimen	An orb-web spider egg sack. First described as "Aranea" columbiae (1878)[49]

Scorpiones

Crustacea

Branchiopoda

Family	Genus	Species	Member	Abundance	Notes	Image
Cyclestheriidae	†Cyclestherioides	†C. wyomingensis	Lake Gosiute (Laney)		A cyclestheriid clam shrimp.[52]
Lynceidae	†Prolynceus	†P. laneyensis	Lake Gosiute (Laney)		A clam shrimp.

Decapods

Family	Genus	Species	Member	Abundance	Notes	Image
Palaemonidae	†Bechleja	†B. rostrata	Fossil Lake		A palaemonid shrimp.[53]
Cambaridae	Procambarus	†P. primaevus	Fossil Lake		A cambarid crayfish.

Ostracoda

Family	Genus	Species	Member	Abundance	Notes	Image
Candonidae	Candona	C. artesensis	Fossil Lake Lake Unita		A candonid ostracod.
		C. rangliensis	Fossil Lake Lake Unita		A candonid ostracod.
		C. whitei	Fossil Lake Lake Gosiute Lake Unita		A candonid ostracod.
	Tuberocypris	T. centronotus	Lake Unita		A candonid ostracod.
†Cyprideidae	†Cypridea	†C. bisulcata	Lake Unita		A cyprideid ostracod.
Cyprididae	†Advenocypris	†A. umbonata	Lake unita		A cypridid ostracod.[54]
	Hemicyprinotus	†H. watsonensis	Lake unita		A cypridid ostracod. One described subspecies H. watsonensis emaciata
	Heterocypris	†H. riflensis	Lake unita		A cypridid ostracod.
		†H. whiteriverensis	Lake unita		A cypridid ostracod.
	Potamocypris	†P. williamsi	Fossil Lake Lake Unita		A cypridid ostracod.
	†Procyprois	†P. ravenridgensis]]	Lake Unita		A cypridid ostracod. Male and female specimens show shell sexual dimorphism
	Pseudoeucypris	†P. pagei	Lake Unita		A cypridid ostracod. First described as Candona pagei (1949) Moved from Cypris pagei (1956)
	Scottia	†S. subquadrata	Lake Unita		A cypridid ostracod.
	Stenocypris	Cf. S. sinuata	Lake Unita		A cypridid ostracod.
Limnocytheridae	Limnocythere	L. sp.	Lake Unita		Indistinct limnocytherid fossils.
	Metacypris	†M. paracordata	Lake Unita		A limnocytherid ostracod.
Notodromadidae	Cyprois	†C. ephraimensis	Fossil Lake Lake Gosiute Lake Unita		A notodromadid ostracod.

Myriapoda

Family	Genus	Species	Member	Abundance	Notes	width=170px	Image
Parajulidae	Julus?			1 specimen	A punitive parajulidae millapede. First described as Iulus telluster (1878). Considered nomen dubium by Hoffman (1969).[55]

Insecta

Coleoptera

Adephaga

Family	Genus	Species	Member	Abundance	Notes	Image
Carabidae	Bembidium	†B. exoletum	Lake Unita		A Bembidiini tribe trechine ground beetle.
	†Carabites	†C. eocenicus	Lake Unita		A ground beetle of uncertain subfamily affiliation.
		†C. exanimus	Lake Uinta		A ground beetle of uncertain subfamily affiliation.[56]
	Harpalus	†H. veterum	Lake Uinta		A Harpalini tribe harpaline ground beetle.
	Lebia	†L. protospiloptera	Lake Unita		A Lebiini tribe harpaline ground beetle.[57]
	†Neothanes	†N. testeus	Lake Gosiute		A ground beetle of uncertain subfamily affiliation.
	Platynus	†P. senex	Lake Gosiute		A Platynini tribe platynine ground beetle.
Dytiscidae	Laccophilus	Indeterminate	Lake Uinta		An indeterminate Laccophilini tribe predaceous diving beetle.

Elateriformia

Family	Genus	Species	Member	Abundance	Notes	Image
Elateridae	Cardiophorus	†C. exhumatus	Lake Unita	1 specimen	A cardiophorine click beetle.[58]
	Ctenicera	†C. velata	Lake Gosiute		A Prosternini tribe dendrometrine click beetle.[59] First named as Corymbites velatus (1876).
	Oxygonus	†O. mortuus	Lake Uinta		A Prosternini tribe dendrometrine click beetle.
Eucnemidae	Epiphanis	†E. deletus	Lake Unita		An Epiphanini tribe melasine false click beetle.

Polyphaga - Bostrichiformia

Family	Genus	Species	Member	Abundance	Notes	Image
Ptinidae	Anobium	†A. deceptum	Lake Gosiute		A Anobiini tribe anobiine death-watch beetle.[60]
		†A. lignitum	Lake Gosiute		A Anobiini tribe anobiine death-watch beetle.
		†A. ovale	Lake Gosiute		A Anobiini tribe anobiine death-watch beetle.
	Stegobium	†S. defunctus	Lake Gosiute		A Stegobiini tribe anobiine death-watch beetle.[61] First described as Sitodrepa defuncta (1876).

Polyphaga - Cucujiformia

Family	Genus	Species	Member	Abundance	Notes	Image
Anthribidae	Euparius	†E. elusus	Lake Gosiute		A Cratoparini tribe anthribine fungus weevil. First named as Cratoparis? elusus (1878)
		†E. repertus	Lake Gosiute		A Cratoparini tribe anthribine fungus weevil. First named as Cratoparis repertus (1878)
	†Pseudochirotenon	†P. eocaenicus	Lake Unita	1 specimen	An Ecelonerini tribe anthribine fungus weevil.[62]
	Trigonorhinus	†T. pristinus	Lake Gosiute		A Trigonorhinini tribe anthribine fungus weevil. First named as Brachytarsus pristinus (1876)
Brentidae	†Palaeonanophyes	†P. zherikhini	Lake Unita		A Nanophyini tribe nanophyine primitive weevil.[63]
	Perapion	†P. rasnitsyni	Lake Uinta		An Aplemonini tribe apionine primitive weevil.
Cerambycidae	Clytus	†C. pervetustus	Lake Uinta		A Clytini tribe cerambycine long-horn beetle.[64] First named as Lema pervetusta (1921)
	Saperda	†S. caroli	Lake Unita		A Saperdini tribe lamiine long-horn beetle.[65]
Chrysomelidae	Bruchus	†B. anilis	Lake Unita		A Bruchini tribe bruchine leaf beetle.
	Cryptocephalus	†C. vetustus	Lake Gosiute		A Cryptocephalini tribe cryptocephaline leaf beetle.
	†Denaeaspis	†D. chelonopsis	Lake Unita		An Imatidiini tribe Cassidine tortoise beetle.[66]
	†Eosacantha	†E. delocranioides	Lake Unita		An Imatidiini tribe Cassidine tortoise beetle.
	†Palaeatalasis	†P. monrosi	Lake Unita	1 specimen	A Megamerini tribe sagrine leaf beetle.[67]
	†Pulchritudo	†P. attenboroughi	Lake Uinta	1 specimen	A sagrine leaf beetle[68]
Cryptophagidae	Antherophagus	†A. priscus	Lake Gosiute		A Cryptophagini tribe cryptophagine silken fungus beetle.
Curculionidae	†Archaeoheilus	†A. deleticius	Lake Uinta		A Molytini tribe molytine weevil. First named as Pachylobius deleticius (1893) Moved from Hylobius deleticius (2015)
		†A. ovalis	Lake Uinta		A Molytini tribe molytine weevil.
		†A. packardii	Lake Gosiute		A Molytini tribe molytine weevil. First named as Hylobius packardii (1893).
		†A. provectus	Lake Gosiute		A Molytini tribe molytine weevil. First named as Hylobius provectus (1876).
		†A. scudderi	Lake Uinta		A Molytini tribe molytine weevil.
	"Coniatus"	†"C." refractus	Lake Uinta		A weevil.[69] Generic placement was deemed dubious by Legalov (2020).[70]
	Cryptorhynchus	†C. annosus	Lake Gosiute		A Cryptorhynchini tribe cryptorhynchine weevil.
	"Dryocoetes"	†"D". carbonarius	Lake Gosiute		A Scolytini tribe scolytine bark beetle. Considered genus and subfamily incertae sedis by Legalov (2015)
	Entimus	†E. primordialis	Lake Uinta		An Entimini tribe entimine weevil.
	Epicaerus	†E. effossus	Lake Gosiute		A Geonemini tribe entimine weevil.[71] First named as Eudiagogus effossus (1876).
		†E. exanimis	Lake Gosiute		A Geonemini tribe entimine weevil. First named as Eudiagogus exanimis (1876).
		†E. saxatilis	Lake Gosiute		A Geonemini tribe entimine weevil. First named as Eudiagogus saxatilis (1876).
		†E. terrosus	Lake Gosiute		A Geonemini tribe entimine weevil. First named as Eudiagogus terrosus (1878).
	Eudiagogus	†E. vossi	Lake Uinta		A Eudiagogini tribe entimine weevil.
	Extinctocentrinus	†E. brevirostris	Lake Uinta		An Apostasimerini tribe conoderine weevil.[72]
	Geraeus	†G. anvilis	Lake Uinta		An Apostasimerini tribe conoderine weevil.
		†G. fossilis	Lake Uinta		An Apostasimerini tribe conoderine weevil.
	Gymnetron	†G. lecontei	Lake Gosiute		A Mecinini tribe curculionine weevil. Considered nomen dubium by Legalov (2015)
	†Lithogeraeus	†L. greenriverensis	Lake Uinta		An Apostasimerini tribe conoderine weevil.
	Nicentrus	†N. curvirostris	Lake Uinta		An Apostasimerini tribe conoderine weevil.
	Ophryastes	†O. compactus	Lake Gosiute		A Tropiphorini tribe entimine weevil. Considered nomen dubium (2011, 2015).[73]
	Otiorhynchus	†O. tumbae	Lake Gosiute		A Otiorhynchini tribe entimine weevil. Considered nomen dubium by Legalov (2015).First named Otiorhynchus dubius (1878)
	†Palaeomallerus	†P. longirostris	Lake Uinta		A Palaeomallerini tribe conoderine weevil.
	"Polygraphus"	†"P". wortheni	Lake Gosiute		A Polygraphini tribe scolytine bark beetle. Considered genus and subfamily incertae sedis by Legalov (2015)
	†Primocentron	†P. wickhami	Lake Uinta		An Eustylini tribe entimine weevil. First described as Otiorhynchus perditus (1876)
	†Pseudophaops	†P. perditus	Lake Gosiute		A Tropiphorini tribe entimine weevil. First named Otiorhynchus perditus (1876)
		†P. wickhami	Lake Uinta		A Tropiphorini tribe entimine weevil.
	†Sciabregma	†S. tenuicornis	Lake Unita		A Sciabregmini tribe molytine weevil.
	"Trypodendron"	†"T". impressus	Lake Gosiute		A Scolytini tribe scolytine bark beetle. Considered genus and subfamily incertae sedis by Legalov (2015) Jr synonym includes Dryocoetes impressus (1878 in paleontology)
Erotylidae	Tritoma	†T. binotata	Lake Gosiute		A Tritomini tribe erotyline pleasing fungus beetle.[74] First named as Mycotretus binotata (1878)
Melandryidae	†Cicindelopsis	†C. eophilus	Lake Unita		A false darkling beetle.
Mordellidae	†Petrimordella	†P. priscula		1 specimen	A tumbling flower beetle morphospecies. First named as Mordella priscula (1925).[75]
		†P. rasnitsyni		1 specimen	A tumbling flower beetle morphospecies. First named as Isotrilophus rasnitsyni (2016).[76]
Nemonychidae	†Eocaenonemonyx	†E. kuscheli	Lake Unita		An Eocaenonemonychini tribe cretonemonychine pine flower weevil.[77]
	†Eugnamptus	†E. decemsatus	Lake Gosiute		An Eocaenonemonychini tribe cretonemonychine pine flower weevil. Considered nomen dubium by Legalov (2013).
		†E. grandaevus	Lake Gosiute		An Eocaenonemonychini tribe cretonemonychine pine flower weevil. First named as Sitona grandaevus (1876) Considered nomen dubium by Legalov (2013).
Peltidae	Indeterminate	Indeterminate	Lake Gosiute		Indeterminate sheild beetle remains. First named as Phenolia incapax.[78]

Polyphaga - Scarabaeiformia

Family	Genus	Species	Member	Abundance	Notes	Image
Scarabaeidae	Aegialia	†A. rupta			An aegialiine scaraboid beetle.[79]
	Phyllophaga	†P. avus	Lake Unita	1 specimen	A melolonthine scaraboid beetle. First described as Melolonthites avus (1921)

Polyphaga - Staphyliniformia

Family	Genus	Species	Member	Abundance	Notes	Image
Hydrophilidae	Berosus	†B. sexstriatus	Lake Gosiute		A Berosini tribe hydrophiline water scavenger beetle.
		†B. tenuis	Lake Gosiute		A Berosini tribe hydrophiline water scavenger beetle.
	Enochrus	†E. primaevus	Lake Gosiute		An enochrine water scavenger beetle. First named as Philhydrus primaevus (1878).
	Hydrobius	†H. decineratus	Lake Gosiute		A Hydrobiusini tribe hydrophiline water scavenger beetle.
	Laccobius	†L. elongatus	Lake Gosiute		A Laccobiini tribe hydrophiline water scavenger beetle.
	Tropisternus	†T. saxialis	Lake Gosiute		A Hydrophilini tribe hydrophiline water scavenger beetle.
		†T. sculptilis	Lake Gosiute		A Hydrophilini tribe hydrophiline water scavenger beetle.
Staphylinidae	Bledius	†B. adamus	Lake Gosiute		A Blediini tribe oxyteline rove beetle.
	Bolitogyrus	Undescribed	Lake Unita		A Staphylinini tribe staphylinine rove beetle.[80]
	Gyrophaena	†G. saxicola	Lake Unita		A Homalotini tribe aleocharine rove beetle.[81]
	Lathrobium	†L. abscessum	Lake Gosiute		A Lathrobiini tribe paederine rove beetle.
	Leistotrophus	†L. patriarchicus	Lake Unita		A Staphylinini tribe staphylinine rove beetle.
	Oxytelus	†O. pristinus	Lake Unita		A Oxytelini tribe oxyteline rove beetle.
	†Staphylinites	†S. obsoletum	Lake Gosiute		A rove beetle of uncertain family affiliations.

Dictyoptera

Family	Genus	Species	Member	Abundance	Notes	Image
Alienopteridae	†Chimaeroblattina	†C. brevipes	Lake Unita		An alienopterid Dictyopteran.[82]
	†Grant	†G. viridifluvius	Lake Unita		An alienopterid Dictyopteran.
Blattidae	Periplaneta	†P.? smithiae	Lake Uinta (Parachute Creek)	1 specimen	A possible periplanetean blattid cockroach.[83]
Blaberidae	Diploptera	†D. gemini	Lake Unita		A Diplopterine beetle cockroache.[84]
		†D. savba	Lake Unita		A Diplopterine beetle cockroache.
		†D. vladimir	Lake Unita		A Diplopterine beetle cockroache.
Ectobiidae	Cariblattoides	†C. labandeirai	Lake Uinta		A Pseudophyllodromiine wood cockroach.[85]
	Ectobius	†E. kohlsi	Lake Unita		An ectobiine Ectobiid cockroach.[86]

Diptera

Family	Genus	Species	Member	Abundance	Notes	Image
Anisopodidae	Mycetobia	†"M." terricola	Lake Gosiute		A wood gnat. First described as Diadocidia? terricola Considered nomen dubium in (2019)[87]
	†Olbiogaster	†O. simplex	Lake Unita		A olbiogastrine wood gnat. First described as Eothereva simplex (1921)
Asilidae	Asilus	†A. palaeolestes	Lake Unita		An Asilini tribe asiline robber fly.
	†Stenocinclis	†S. anomala	Lake Gosiute		A dasypogonine robber fly.
Anthomyiidae	Anthomyia	†"A." winchesteri	Lake Unita		First described as an anthomyiid fly. Deemed nomen dubium without evidence in 1996.[88]
Bibionidae	Plecia	†P. akerionana	Lake Unita (Parachute Creek Member)		A bibonid marchfly. The smallest species from the formation.[89]
		†P. dejecta			A bibonid marchfly.
		†P. pealei	Fossil Lake (Fossil Butte Member)	Most common insect in Fossil Lake	A bibonid marchfly.
		†P. rhodopterina	Lake Unita		A bibonid marchfly.
		†P. winchesteri	Lake Unita		A bibonid marchfly.
		†P. woodruffi	Lake Unita		A bibonid marchfly.
Blephariceridae	†Philorites	†P. pallescens	Lake Unita		A net-winged midge of uncertain subfamily affinities.
Cecidomyiidae	Lasioptera	†L. recessa	Lake Unita		A Lasiopterini tribe cecidomyiine gall midge.
	†Lithomyza	†L. condita	Lake Unita		A Lestremiini tribe Lestremiine gall midge.
Chaoboridae	"Corethra"?	†"C".? exita	Lake Unita		A punitive phantom midge.
Chironomidae	Chironomus	†C. depletus	Lake Unita		A Chironomini tribe chironomine nonbiting midge.
		†C. patens	Lake Unita		A Chironomini tribe chironomine nonbiting midge.
	Undescribed	Undescribed	Lake Unita	Numerous	undescribed midge larvae.
Culicidae	Culex	†C. winchesteri	Lake Unita		A Culicini tribe culicine mosquito.[90]
Cylindrotomidae	Cylindrotoma	†C. veterana	Lake Unita		A cylindrotomine long bodied cranefly.
	†Cyttaromyia	†C. fenestrata	Lake Unita	1 specimen	A cylindrotomine long bodied cranefly species.[91] [92]
		†C. freiwaldi	Lake Unita	2 specimens	A cylindrotomine long bodied cranefly species.
		†C. fuscula	Lake Unita	1 specimen	A cylindrotomine long bodied cranefly species.First described as Asilopsis fusculus (1921).
		†C. gelhausi	Lake Unita	3 specimens	A cylindrotomine long bodied cranefly species.
		†C. obdurescens	Lake Unita	1 specimen	A cylindrotomine long bodied cranefly species.
		†C. reclusa	Lake Unita	1 specimen	A cylindrotomine long bodied cranefly species.
Empididae	†Protoedalea	†P. brachystoma	Lake Uinta		A balloon fly of uncertain subfamily affinity.
	Rhamphomyia	†R. enena	Lake Unita		An Empidini tribe empidine balloon fly.
Heleomyzidae	Heteromyza	†H. detecta	Lake Unita		A heleomyzine Heleomyzid fly.
Hippoboscidae	†Eornithoica	†E. grimaldii	Fossil Lake (Fossil Butte Member)	1 specimen	A Hippoboscoidea fly Pupipara.[93]
Keroplatidae	Palaeoplatyura	†P. eocenica	Lake Uinta		An Orfeliini tribe keroplatine predatory fungus gnat.
Limoniidae	Dicranomyia	†D. primitiva	Lake Unita		A limoniine limoniid cranefly.
		†D. rostrata	Lake Unita		A limoniine limoniid cranefly. Includes Tipula decrepita (1877).[94]
		†D. stigmosa	Lake Unita		A limoniine limoniid cranefly. Includes Tipula tecta (1877).
	†Pronophlebia	†P. rediviva	Lake Unita		A limoniid cranefly of uncertain subfamily placement.
	Rhabdomastix	†R. scudderi	Lake Unita		A chioneine limoniid cranefly.[95] First named as Gonomyia scudderi (1921).
Mycetophilidae	Gnoriste	†G. dentoni	Lake Unita		A gnoristine fungus gnat.
	Leptomorphus	†L. palaeospilus	Lake Unita		A sciophiline fungus gnat.[96] First named as Diomonus palaeospilus.
	Mycetophila	†M. occultata	Lake Unita		A Mycetophilini tribe Mycetophiline fungus gnat..
	†Sackenia	†S. arcuata	Lake Unita		A Mycetophiline fungus gnat.
		†S. gibbosa	Lake Gosiute		A Mycetophiline fungus gnat.[97]
	Undescribed	Undescribed	Lake Unita	Numerous	undescribed larvae.
Mythicomyiidae	Acrocera	†A. hirsuta	Lake Unita		An acrocerine mythicomyiid fly. Possibly nomen dubium
	Reissa	†R. kohlsi			A mythicomyiine mythicomyiid fly[98]
Oestridae	Cuterebra	†C. ascarides	Lake Unita	Numerous	A cuterebrine bot fly species. First named as Musca ascarides (1877) Moved from Lithohypoderma ascarides (1916)
		†C. bibosa	Lake Unita	Numerous	A cuterebrine bot fly species. First named as Musca bibosa (1877)
	Dermatobia	†D. hydropica	Lake Unita		A cuterebrine bot fly.
Stratiomyidae	Nemotelus	†N. eocenicus	Lake Unita		A soldier fly.
	Odontomyia	Undescribed	Lake Unita	Numerous	undescribed soldier fly larvae.
	Sargus	†S. vetus	Lake Unita		A soldier fly.
Psychodidae	Indeterminate	Indeterminate	Lake Untia		An indeterminate psychodid fly.[99] First described as Culex proavitus (1877).
Sciomyzidae	†Sciomyza?	†S.? manca	Lake Gosiute Lake Unita		A Sciomyzini tribe sciomyzine marsh fly.
		†S.? disjecta	Lake Gosiute		A Sciomyzini tribe sciomyzine marsh fly.
Syrphidae	†Asarkina	†A. quadrata	Lake Gosiute		A Syrphini tribe syrphine hoverfly. First described as Milesia quadrata (1878)
	Cheilosia	†C. ampla	Lake Gosiute		A Rhingiini tribe eristaline hoverfly.
	Eristalis	†E. lapideus			An Eristalini tribe eristaline hoverfly.
	Syrphus	†S. eocenicus	Lake Uinta		A Syrphini tribe syrphine hoverfly.
		†S. lithaphidis	Lake Unita		A Syrphini tribe syrphine hoverfly.
	Undescribed	Undescribed	Lake Unita	Numerous	undescribed hoverfly larvae.
Tabanidae	Undescribed	Undescribed	Lake Unita	Numerous	undescribed larvae.
Tipulidae	†Spiladomyia	†S. simplex	Lake Unita		A tipulid cranefly of uncertain subfamily affinities.
	Tipula	†T. americana	Lake Unita		A tipuline tipulid cranefly.[100]
Incertae sedis	†Acanthomyites	†A. aldrichi	Lake Unita		A brachyceran fly of uncertain family affinities. First treated as a house fly.
	†Poliomyia	†P. recta	Lake Gosiute		A schizophoran fly of uncertain affinity.[101]
	†Vinculomusca	†V. vinculata	Lake Unita		A fly pupal case morphospecies of uncertain affinity.[102] First named as Musca vinculata (1877)

Hemiptera

Family	Genus	Species	Member	Abundance	Notes	Image
Aphididae	†?Siphonophoroides	†?S. antiqua	Lake Gosiute		A possible drepanosiphine aphid. First identified as possibly Lithaphis diruta (1890)
Cercopidae	Cercopis	†C. cephalinus	Lake Unita		A Cercopini tribe cercopine froghopper.
Cicadellidae	Aphrodes	†A. adae	Lake Gosiute		An Aphrodini tribe aphrodine leafhopper. First named as Acocephalus adae (1878).
	Cicadella	†C. parobtecta	Lake Unita		A Cicadellini tribe cicadelline leafhopper. A replacement name for Tettigonia obtecta (1877 non-Fabricius, 1803)[103]
		†C. scudderi	Lake Unita		A Cicadellini tribe cicadelline leafhopper.
	Coelidia	†C. wyomingensis	Fossil Lake (Fossil Butte Member)	1 specimen	A coelidiine leafhopper.
	Iassus	†I. lapidescens	Lake Uinta		An Iassini tribe iassine leafhopper.[104] First named as Bythoscopus lapidescens (1877)
	Thamnotettix	†T. packardi	Lake Unita		An Athysanini tribe deltocephaline leafhopper. First named as Erythroneura eocenica (1921)
Cixiidae	Cixius	†C.? hesperidum	Lake Gosiute		A possible Cixiini tribe cixiine cixiid planthopper. Considered nomen dubium by Swedo and Lefebvre (2004).[105]
	†Oliarites	†O. terrentula	Lake Gosiute		A cixiine cixiid planthopper. First named as Mnemosyne terrentula (1878) Considered nomen dubium by Swedo and Lefebvre (2004).[106]
Cydnidae	Cyrtomenus	†C. concinnus	Lake Gosiute		A Geotomini tribe cydnine burrowing bug.
	†Procydnus	†P. mamillanus	Lake Gosiute		A cydnine burrowing bug. First named as Cydnus? mamillanus (1878).
	†Stenopelta	↑S. punctulata	Lake Gosiute		A cydnine burrowing bug. First named as Aethus punctulatus (1878).
Delphacidae	Delphax	†D. veterum	Lake Uinta		A Delphacini tribe delphacine delphacid froghopper
Dinidoridae	†Dinidorites	†D. margiformis	Lake Unita		A dinidorid shield bug of uncertain subfamily affinities.
Fulgoridae	Aphaena	†A. atava	Lake Unita		A lanternfly of uncertain subfamily affiliations First named as Aphana atava (1877)
		†A. rotundipennis	Lake Gosiute		A lanternfly of uncertain subfamily affiliations First named as Aphana rotundipennis (1878)
	Fulgora	†F. granulosa	Lake Gosiute		A Fulgorini tribe fulgorine lanternfly.
		†F. populata	Lake Gosiute		A Fulgorini tribe fulgorine lanternfly.
	Lystra?	†L.? leei	Lake Gosiute		A Lystrini tribe poiocerine lanternfly.
		†L.? richardsoni	Lake Gosiute		A Lystrini tribe poiocerine lanternfly.
Gerridae	†Telmatrechus	†T. parallelus	Fossil Lake (Fossil Butte Member) Lake Uinta (Laney Member)		A water strider.[107]
Lophopidae	†Scoparidea	†S. nebulosa	Lake Unita		A lophopid planthopper.
Lyctocoridae	Lyctocoris	†L. terreus	Lake Gosiute		A Lyctocorini tribe lyctocorine Lyctocorid bug. First named as Rhyparochromus? terreus (1878)
Nogodinidae	†Detyopsis	†D. scudderi	Lake Unita		A nogodinid planthopper.
		†D. scudderi	Lake Unita		A nogodinid planthopper.
	†Lithopsis	†L. delicata	Lake Unita		A nogodinid planthopper.
		†L. fimbriata	Lake Gosiute		A nogodinid planthopper.
		†L. simillima	Lake Unita		A nogodinid planthopper.
	"Paralatindia"	†"P." saussurei	Lake Gosiute		A nogodinid planthopper. First described as a Paralatindia genus cockroach. Considered nomen dubium by Shcherbakov (2006)[108]
Reduviidae	†Aphelicophontes	†A. danjuddi	Lake Unita		A Harpactorini tribe Harpactorine Assassin bug.[109]
Rhopalidae	Corizus	†C. guttatus	Lake Gosiute		A Rhopalini tribe rhopaline scentless plant bug. First named as Reduvius guttatus (1878).
Rhyparochromidae	Aphanus	†A. petrensis	Lake Unita		A Gonianotini tribe rhyparochromine seed bug. First named as Pachymerus petrensis (1877).[110]
Ricaniidae	†Dilaropsis	†D. ornatus	Lake Unita		A Ricaniid plant hopper.
	†Hammapteryx?	†H.? ceryniiformis	Lake Unita		A Ricaniid plant hopper.
		†H.? lepidoides	Lake Unita		A Ricaniid plant hopper.
Tingidae	†Gyaclavator	†G. kohlsi	Lake Unita		A lace bug of uncertain subfamily affiliation.[111]	center|upright|frameless
Incertae sedis	"Delphax"	†"D." senilis			First described as an delphacine froghopper. Considered nomen dubium
	†Diaplegma	†D. obdormitum	Lake Gosiute		A fulgoroid leafhopper of uncertain familial affinities. First placed as a cixiid planthopper.
	†Emiliana	†E. alexandri	Lake Unita		A fulgoroid leafhopper of uncertain familial affinities. First placed as a Tropiduchid froghopper
	†Eoliarus	†E. lutensis	Lake Gosiute		A fulgoroid leafhopper of uncertain familial affinities. First described as Oliarus lutensis (1890)
		†E. quadristictus			A fulgoroid leafhopper of uncertain familial affinities.
	†Protoliarus	†P. humatus	Lake Uinta		A fulgoroid leafhopper of uncertain familial affinities. First described as a lanternfly.

Hymenoptera

Family	Genus	Species	Member	Abundance	Notes	Image
Apidae	†Euglossopteryx	†E. biesmeijeri	Lake Unita (Parachute Creek Member)		An apid bee.[112]
Bethylidae	Eupsenella	E. sp.			An old world Bethylidae wasp.[113]
Braconidae	Bracon	†B. laminarum	Lake Gosiute		A Braconini tribe braconine braconid wasp.
	†Colemanus	†C. keeleyorum	Lake Unita (Parachute Creek Member)		A Proteropini tribe ichneutine braconid wasp.[114]
	†Eobracon	†E. cladurus	Lake Unita		A braconid wasp of uncertain subfamily affinities.
	Hormiopterus	†H. petrinus	Lake Unita		A braconid wasp of uncertain subfamily affinities. First named as Ichneumon petrinus[115]
Eurytomidae	Eurytoma	†E. antiqua	Lake Gosiute		An eurytomine Eurytomid chalcid wasp. First described as Decatoma antiqua (1878).
Formicidae	†Archimyrmex	†A. rostratus	Lake Unita	Numerous	A myrmeciine ant.
	†Camponotites	†C. vetus	Green River Formation?	1 specimen	A formicine ant of uncertain affiliation. First described as Camponotus vetus (1877)[116] The type locality was not specified by Scudder, and may Be G.R.F. or White River Formation.[117]
	Dolichoderus	†D. kohlsi	Lake Uinta	Numerous	A Dolichoderus species ant.[118]
	†Eoformica	†E. globularis	Lake Unita	2 specimens	An ant of unidentified affiliation.
		†E. magna	Lake Unita	1 specimen	An ant of unidentified affiliation.
		†E. pinguis	Lake Gosiute Lake Uinta	Numerous	An ant of unidentified affiliation. Moved from Liometopum pingue (1877) Includes the jr synonyms Lasius terreus (1878) and Eoformica eocenica (1921)
		Indeterminate	Lake Unita	2 specimen	Two partial specimens that may be referable to Eoformica.
	†Kohlsimyrma	†K. gracilis	Lake Unita	2 specimens	An ant of uncertain affiliation.
		†K. laticeps	Lake Unita	1 specimen	An ant of uncertain affiliation.
		†K. longiceps	Lake Unita	3 specimens	An ant of uncertain affiliation.
	†Mianeuretus	†M. eocenicus	Lake Unita	2 specimens	An aneuretine ant.
		Indeterminate	Lake Unita	1 specimen	A partial specimen that may belong to Mianeuretus.
	†Myrmecites	†M. rotundiceps	Lake Unita	1 specimen	A myrmicine ant of uncertain affiliation.
	Pachycondyla	†P. labandeirai	Lake Gosiute	1 specimen	A Pachycondyla species ant.
	†Ponerites	†P. coloradensis	Lake Unita	1 specimen	A ponerine ant of uncertain affiliation.
		†P. eocenicus	Lake Unita	1 specimen	A ponerine ant of uncertain affiliation.
		†P. hypoponeroides	Lake Unita	1 specimen	A ponerine ant of uncertain affiliation.
	†Proiridomyrmex	†P. vetulus	Lake Unita	1 specimen	A dolichoderine of uncertain affinity.
		Indeterminate	Lake Unita	1 specimen	A partial specimen that may belong to Proiridomyrmex.
	†Solenopsites	†S. minutus	Lake Unita	1 specimen	A Solenopsis like ant of uncertain affiliation.
	†Titanomyrma	†T. lubei	Fossil Lake (Angelo Member)	3 specimens	A titan ant species.[119] [120]
	Indeterminate	Indeterminate	Lake Unita	1 specimen	A partial ant specimen that may belong to Formica.
	Indeterminate	Indeterminate	Lake Unita	3 specimens	Partial ant specimens of unidentified subfamily placement.
Ichneumonidae	Allomacrus	†A. lanhami	Lake Unita		A cylloceriine darwin wasp.[121] First named as Plectiscidea lanhami (1941)[122]
	†Carinibus	†C. molestus	Lake Unita		A darwin wasp of uncertain subfamily placement.[123]
	Eclytus?	†E.? lutatus	Lake Gosiute?		A possible Eclytini tribe tryphonine darwin wasp.
	†Eopimpla	†E. grandis	Lake Unita		A darwin wasp of uncertain subfamily placement.
	Glypta?	†G.? transversalis	Lake Gosiute?		A Glyptini tribe banchine darwin wasp.
	†Ichninsum	†I. appendicrassum	Lake Unita		A pimpline darwin wasp.
	†Lithotorus	†L. cressoni]	Lake Gosiute?		An Orthocentrine darwin wasp.
	Mesoclistus?	†M.? yamataroti	Lake Unita		A possible Coleocentrini tribe acaenitine darwin wasp.
	Pimpla?	†P.? eocenica	Lake Unita		A possible Pimplini tribe pimpline darwin wasp.[124]
	Phygadeuon	†P. petrifactellus	Lake Uinta		A Phygadeuontini tribe phygadeuontine darwin wasp.
	Rhyssa?	†R.? juvenis	Lake Gosiute?		A Rhyssini tribe rhyssine darwin wasp.
	Scambus?	†S.? mandibularis	Lake Unita		A possible Ephialtini tribe pimpline darwin wasp.
		†S.? parachuti	Lake Unita		A possible Ephialtini tribe pimpline darwin wasp.
	†Tilgidopsis	†T. haesitans	Lake Unita		A darwin wasp of uncertain subfamily placement.
	†Trymectus	†T. amasidis	lake Unita		A darwin wasp of uncertain subfamily placement. First described as Tryphon amasidis (1931)
Siricidae	†Eoteredon	†E. lacoi	Fossil Lake	1 specimen	A siricid Horntail wood-wasp.[125]
Tenthredinidae	Eutomostethus	†E. karimae	Lake Unita		A blennocampine common sawfly.[126]
Xyelidae	†Proxyelia	†P. pankowskii	Fossil Lake	1 specimen	A xyelid sawfly[127]

Neuroptera

Family	Genus	Species	Member	Abundance	Notes	Image
Berothidae	†Xenoberotha	†X. angustialata	Lake Unita (Parachute Creek Member)	1 specimen	A berothine beaded lacewing.[131]
Hemerobiidae	Megalomus?	†M.? coloradensis	Lake Unita		A possible Megalomus brown lacewing.[132]
	†Pseudonotherobius	†P. kohlsi	Lake Unita		A brown lacewing.
Mantispidae	†Protonolima	†P. mantispinoformis	Lake Unita	1 specimen	A calomantispine mantidfly.[133]
Myrmeleontidae	†Epignopholeon	†E. sophiae	Lake Unita (Parachute Creek Member)	1 specimen	A probable Gnopholeontini tribe myrmeleontine antlion.[134]
Osmylidae	†Osmylidia	†O. taliae	Lake Unita	1 specimen	A protosmyline Osmylid lance lacewing[135]
Incertae sedis	†Callospilopteron	†C. ocellatum			A lacewing of uncertain affiliation. First identified as an hemipteran planthopper.

Odonata

Primarily based on Bechly et al (2020):^[138]

Family	Genus	Species	Member	Abundance	Notes	Image
Amphipterygidae	†Protamphipteryx	P. basalis	Fossil Lake		An amphipterygine amphipterygid damselfly.[139]
†Carleidae	†Carlea	C. eocenica	Fossil Lake	1 specimen	A carleid damselfly.
Cordulibellulidae	†Cordulibellula	C. inopinata	Lake Uinta (Parachute Creek)		A cordulibellulid dragonfly the last member of an ancient lineage.[140]
Corduliidae	†Stenogomphus	†S. carletoni	Lake Unita		A green-eyed skimmer.
		†"S." scudderi	Lake Unita		First described as a green-eyed skimmer. Deemed nomen dubium in 1994.[141]
Dysagrionidae	†Dysagrion	D. fredericii	Lake Gosiute (Laney Member)	16 specimens	A dysagrionine dysagrionid damselfly.
		D. integrum	Fossil Lake (Fossil Butte Member)	1 specimen
		D. lakesii	Lake Gosiute (Laney Member)	1 specimen
		D. packardii	Lake Gosiute (Laney Member)	1 specimen
	†Petrolestes	†P. hendersoni	Lake Uinta (Parachute Creek Member)	1 specimen	A petrolestine dysagrionid damselfly.
		†P. inexpectatus	Lake Uinta	1 specimen	A petrolestine dysagrionid damselfly.
	†Tenebragrion	†T. tynskyi	Fossil Lake		A dysagrionid damselfly.[142]
		†T. shermani	Fossil Lake		A dysagrionid damselfly.
Calopterygidae	†Eocalopteryx	E. atavina	Lake Gosiute	1 specimen	A calopterygid damsefly.
Euphaeidae	†Eodysphaea	E. magnifica	Lake Uinta (Parachute Creek)		A euphaeid damselfly.
	†Epallagites	E. avus	Lake Unita
	†Labandeiraia	L. americaborealis	Lake Uinta (Parachute Creek)	1 specimen
		L. riveri	Fossil Lake	1 specimen
	†Litheuphaea	L. coloradensis	Lake Uinta (Parachute Creek)	1 specimen
†Eolestidae	†Eolestes	E. syntheticus	Lake Unita		An eolestid damselfly.[143]
Megapodagrionidae	†Viridiflumineagrion	V. aasei	Lake Uinta (Parachute Creek Member)	1 specimen	A "megapodagrionid" damselfly.[144]
Thaumatoneuridae	†Gusagrion	†G. coloratum	Lake Uinta	1 specimen	An eodysagrionine thaumatoneurid damselfly.[145]
	†Oreodysagrion	O. tenebris	Fossil Lake		A thaumatoneurid damselfly.
	†Tynskysagrion	T. brookeae	Fossil Lake		A thaumatoneurid damselfly.
†Urolibellulidae	†Kohlslibellula	K. lini	Lake Uinta (Parachute Creek)		A urolibellulid dragonfly
	†Urolibellula	U. eocaenica	Lake Uinta (Parachute Creek)		A urolibellulid dragonfly
Zacallitidae	†Zacallites	Z. balli			A zacallitid damselfly.
		Z. cockerelli	Fossil Lake
incertae sedis	†Eopodagrion	"E." abortivum	Lake Gosiute (Laney Member)	1 specimen	A damselfly of uncertain affinities. First named as Podagrion abortivum (1878) Deemed nomen dubium by Nel (2022).[146]
		E. scudderi	Lake Gosiute (Laney Member)	3 specimens	A damselfly of uncertain affinities.

Orthoptera

Family	Genus	Species	Member	Abundance	Notes	Image
Eumastacidae	†Eoerianthus	†E. eocaenicus	Lake Unita		A monkey grasshopper.[147]
		†E. multispinosus	Lake Gosiute		A monkey grasshopper. First described as Tyrbula multispinosa (1890.
Gryllidae	Eogryllus	†E. unicolor	Lake Unita		A Gryllini tribe grylline true cricket.
	Eogryllus?	†E.? elongatus	Lake Unita		A Gryllini tribe grylline true cricket. Tentatively placed in Eogryllus
	†Pronemobius	†P. induratus	Lake Gosiute		A Gryllini tribe grylline true cricket.
		†P. ornatipes	Lake Unita		A Gryllini tribe grylline true cricket.
		†P. tertiarius	Lake Gosiute Lake Unita		A Gryllini tribe grylline true cricket. Tentatively placed in Pronemobius First described as Nemobius tertiarius (1878).[148]
Gryllotalpidae	†Pterotriamescaptor?	†P.? americanus	Lake Unita		A gryllotalpine mole cricket.
Mogoplistidae	†Eomogoplistes	†E. longipennis	Lake Unita		A Mogoplistini tribe Mogoplistine scaly cricket.
	†Pteromogoplistes	†P. grandis	Lake Unita		A Mogoplistini tribe Mogoplistine scaly cricket.
		Indeterminate	Lake Unita		Indeterminate scaly cricket fossils.
	†Pteromogoplistes?	†P.? smithii	Lake Unita		A Mogoplistini tribe Mogoplistine scaly cricket. Tentatively moved from Pronemobius smithii (1890)
Phalangopsidae	Eotrella	†E. mira	Lake Unita		A phalangopsine spider cricket.
	Eozacla	†E. arachnomorpha	Lake Unita		A phalangopsine spider cricket.
		†E. problematica	Lake Unita		A phalangopsine spider cricket.
Tetrigidae	Eotetrix	†E. unicornis	Lake Unita		A batrachideine pygmy grasshopper.
Tettigoniidae	Arethaea	†A. solterae	Lake Uinta (Parachute Creek)	1 specimen	A roundhead katydid.[149]
Trigonidiidae	Indeterminate	Indeterminate	Lake Unita		Indetermiante trigonidiine sword-tail cricket remains.
Incertae sedis	†Tettoraptor	†T. maculatus	Lake Unita		A Tettigoniid grasshopper of uncertain affiliations.

Strepsiptera

Family	Genus	Species	Member	Abundance	Notes	Image
Myrmecolacidae	Caenocholax	†C. barkleyi	Lake Unita (Parachute Creek Member)	1 specimen	A male twisted-winged insect.[150]
		†C. palusaxus	Lake Unita (Parachute Creek Member)	1 specimen	A male twisted-winged insect.[151]

Thysanoptera

Family	Genus	Species	Member	Abundance	Notes	Image
Aeolothripidae	†Lithadothrips	†L. vetustus	Lake Uinta		An Aeolothripid thrips. First named as Lithadothrips vetusta[152]
	†Palaeothrips	†P. fossilis	Lake Unita		An Aeolothripid thrips.[153]
Melanthripidae	Melanothrips	†M. extincta	Lake Unita		A melanthripid thrips.

Trichoptera

Family	Genus	Species	Member	Abundance	Notes	Image
Hydropsychidae	Hydropsyche	†H. operta	Lake Unita		A hydropsychine net-spinning caddisfly. First named as Phryganea operta (1877)
Hydroptilidae	Hydroptila	†H. phileos	Lake Unita		A Hydroptilini tribe hydroptiline purse-case caddisfly.
Limnephilidae	†Indusia	†I. calculosa	Lake Gosiute?		A northern caddisfly larval case morphospecies.
	Limnephilus	†L. eocenicus	Lake Unita	1 specimen	A northern caddisfly species.
Incertae sedis	†Litholimnephilops	†L. yinani	Lake Unita		A caddis fly of uncertain affiliations.[154]

Mollusks

Based on Grande (1984):^{[155] [156]}

Bivalvia

Family	Genus	Species	Member	Abundance	Notes	Image
Sphaeriidae	Sphaerium	S. sp.	Lake Gosiute		A fingernail clam.
Unionidae	Plesielliptio	P. priscus			A unionid river mussel.
		P. sp. "A"

Gastropoda

Family	Genus	Species	Member	Abundance	Notes	Image
Bulimulidae	Oreoconus	O. sp.	?		An undescribed bulimulid land snail.
Gastrocoptidae	Gastrocopta	G. sp.			An undescribed gastrocoptid land snail.
Hydrobiidae	Hydrobia		Lake Gosiute (Laney, Tipton Shale, Wilkins Peak)		An undescribed mud snail. Affiliated to Hydrobia utahensis
		H. sp.	Lake Gosiute (Tipton Shale)		An undescribed mud snail.
Lymnaeidae	Lymnaea	L. similis	Lake Gosiute (Laney)		A pond snail.
		L. sp.	Lake Gosiute (Tipton Shale)		A undescribed pond snail.
	†Pleurolimnaea	P. tenuicosta	Lake Gosiute (Tipton Shale?)		A pond snail. First named Limnaea tenuicosta (1856)
Planorbidae	Biomphalaria	B. aequalis	Lake Gosiute (Laney, Tipton Shale)		A ramshorn snail.
		B. storchi	Lake Gosiute (Laney?, Tipton Shale)		A ramshorn snail.
	†Headonia	†H. pseudoammonius	Lake Gosiute (Laney, Tipton Shale)		A ramshorn snail First reported as Biomphalaria pseudoammonius (1820)
	?Drepanotrema	?D. sp.	Lake Gosiute (Laney)		An undescribed ramshorn snail.
	Gyraulus	G. militaris	Lake Gosiute (Tipton Shale)		A ramshorn snail.
	Omalodiscus	†O. cirrus	Lake Gosiute (Tipton Shale)		A ramshorn snail.
Physidae	Physa	†?P. longiuscula	Lake Gosiute (Tipton Shale)		An bladder snail.
		P. sp.	Lake Gosiute (Laney)		An undescribed bladder snail.
	Physella	†P. bridgerensis	Lake Gosiute (Laney, Tipton Shale)		A bladder snail. First named Physa bridgerensis (1873)
		†P. pleromatis	Lake Gosiute (Tipton Shale)		A bladder snail. First named Physa pleromatis (1877)
Pleuroceridae	Elimia	†E. tenera	Lake Gosiute (Laney, Tipton Shale, Wilkins Peak)		A pleurocerid snail. the most common snail in the formation The source of the famous "Turritella agate".
Urocoptidae	Holospira	H. sp.			An undescribed urocoptid land snail.
Valvatidae	Valvata	†V. subumbilicata	Lake Gosiute (Laney, Tipton Shale)		A valve snail.
		Cf. †V. filosa			An undescribed valve snail. Similar to Valvata filosa
Viviparidae	Viviparus	†V. meeki	Lake Gosiute (Laney, Tipton Shale)		A river snail. First named Paludina trochiformis (1856 non-von Zieten, 1830)
		†V. paludinaeformis	Lake Gosiute (Tipton Shale)		A river snail.

Fungi

Genus	Species	Member	Abundance	Notes	Image
†Caenomyces	C. eucalypti	Lake Unita	1 specimen	A possible ascomycotan leaf spot fungus morphospecies. Hosted on a Syzygioides americana leaf.[157] [158] Brown (1934) considered all leaf spots of the formation to be indeterminate †Sphaerites species.[159]
	†C. planerae		1 specimen	A possible ascomycotan leaf spot fungus morphospecies. Hosted on a Cedrelospermum nervosum leaf.[160] [161] Brown (1934) considered all leaf spots of the formation to be indeterminate †Sphaerites species.
	†C. sapindicola		1 specimen	A possible ascomycotan leaf spot fungus morphospecies. Hosted on the Toxicodendron winchesteri holotype leaf.[162] Brown (1934) considered all leaf spots of the formation to be indeterminate †Sphaerites species.
†Eoglobella	†E. longipes	Lake Unita	1 specimen	A Staurosporaen fungal spore of uncertain affiliation.[163]
Entophlyctis	†E. willoughbyi	Lake Gosiute (Wilkins Peak Member)		An aquatic chytridialean fungi.[164]
Pleotrachelus	†P. askaulos	Lake Gosiute (Wilkins Peak Member)		An aquatic chytridialean fungi.
†Sphaerites	†S. myricae	Lake Gosiute	1 specimen	A possible ascomycotan leaf spot fungus morphospecies. Hosted on a "Rhus" nigricans leaf. First described as Sphaeria myricae (1878)[165] Brown (1934) considered all leaf spots of the formation to be indeterminate †Sphaerites species.

Unicellular microbiota

Cyanobacteria

Family	Genus	Species	Member	Abundance	Notes	width=170px	Image
Chroococcaceae	Chroococcus	?Cf. C. dispersus	Lake Unita		An aquatic chroococcacean cyanobacteria.[166]
		Cf. C. westii	Lake Unita		An aquatic chroococcacean cyanobacteria.
Coleofasciculaceae	Stigonema	†S. anchistina	Lake Gosiute (Wilkins Peak Member)		An aquatic coleofasciculacean cyanobacteria.[167]
	Symploca	†S. hedraia	Lake Gosiute (Wilkins Peak Member)		An aquatic coleofasciculacean cyanobacteria.
Gomontiellaceae	Crinalium	?Cf. C. endophyticum	Lake Unita		An aquatic gomontiellaceous cyanobacteria.
		C. sp.	Lake Unita		An aquatic gomontiellaceous cyanobacteria. First identified as Spirulina sp. (1916)
Hapalosiphonaceae	Hapalosiphon	H. confervaceus	Lake Unita		An aquatic hapalosiphonaceous cyanobacteria.

Algae

Family	Genus	Species	Member	Abundance	Notes	Image
Chaetophoraceae	Stigeoclonium?		Lake Unita	1 specimen	A possible chaetophoraceous chlorophyte green algae.
Chlorellaceae	†Chlorellopsis	†C. coloniata			A unicellular stromatolite forming green algae.[168]
	†Confervites	†C. mantiensis	Lake Uinta (Manti beds)		A filamentous stromatolite forming green algae.
Hydrodictyaceae	Tetraedron		Lake Unita	1 specimen	A hydrodictyaceous chlorophyte green algae.
Microsporaceae	Microspora	Cf. M. pachyderma	Lake Unita	1 specimen	A sphaeroplealean chlorophyte green algae.
Oocystaceae	Cf. Eremosphaera	E. sp.	Lake Unita		An oocystacean chlorophyte green algae.
Scenedesmaceae	Coelastrum	Cf. C. verrucosum	Lake Unita		A sphaeroplealean chlorophyte green algae.
Schizochlamydaceae	Schizochlamys	†S. haywellensis	Lake Gosiute (Wilkins Peak Member)		A sphaeroplealean chlorophyte green algae.

Amoebozoans

Family	Genus	Species	Member	Abundance	Notes	Image
Difflugiidae	Difflugia	†D. calcifera	Lake Unita	1 specimen	A difflugiid lobose testate amoeba.
Quadrulidae	Quadrula	†Q. minuta	Lake Unita	1 specimen	A quadrulid lobose testate amoeba.

Cercozoans

Plants

Ferns

Family	Genus	Species	Member	Abundance	Notes	Image
Aspleniaceae	Asplenium	†A. delicatula			A spleenwort fern leaf morphospecies.[171] First described as Aneimia delicatula (1929) Includes Phyllites furcivena (1934) Jr homonym of Asplenium delicatulum
		†A. eoligniticum	Lake Unita	2 specimens	A spleenwort fern leaf morphospecies.
Equisetaceae	Equisetum	†E. haydenii	Not Green River Formation		A scouring rush morphospecies. First ascribed to the formation Excluded by Knowlton (1923) as a Washakie Formation species.
		†E. winchesteri	Lake Gosiute	1 specimen	A scouring rush morphospecies.
		†E. wyomingense	Lake Gosiute Lake Unita		A scouring rush morphospecies.
Lygodiaceae	Lygodium	†L. dentoni	Rejected	1? specimen	A climbing fern species. Type specimens missing by 1923 Brown (1934) rejected per Knowlton and dubious locality.
		†L. kaulfussi	Fossil Lake Lake Gosiute Lake Unita	Numerous	A climbing fern species. First described as Lygodium neuropteroides (1870)
Marattiaceae	Danaea	†D. coloradensis	Lake Unita	1 specimen	A possible marattiaceous fern leaf morphospecies. The genus affiliation was questioned by MacGinitie (1969).
Marsileaceae	Marsilea	†M. sprungerorum	Lake Uinta (Parachute Creek Member)	Numerous	A water clover fern.[172]
Osmundaceae	Osmunda?	†Q.? sepulta	Lake Gosiute	2 specimens	A possible osmundaceous fern leaf morphospecies. First described as Pecopteris sepulta (1883)
Salviniaceae	Azolla	†A. berryi	Lake Unita	1 specimen	A mosquito fern morphospecies.
Pteridaceae	Acrostichum	†A. hesperium	Lake Gosiute	3 specimens	A parkerioid leather fern leaf morphospecies.[173]

Cycadalean palynomorphs

Genus	Species	Member	Abundance	Notes	width=170px	Image
†Cycadopites	†C. sp.	Lake Uinta (Parachute Creek Member)	Numerous	Various Cycas - Zamia Cycad palynomorphs.
†Dioonipites	†D. sp.		5 specimens	A Dioon like Cycad palynomorph.

Conifers

Family	Genus	Species	Member	Abundance	Notes	Image
Cupressaceae	Sequoia	Cf. †S. affinis	Lake Unita		A coast redwood relative.
Rowspan=3	Pinaceae	Picea	†P. pinifructus	Lake Unita	1 specimen	First identified as a spruce seed. Deemed a jr synonym of Cercidiphyllum arcticum by Brown (1939).[174]
Pinus	†P. balli	Lake Unita	1 specimen	A 3-needle pine morphospecies.
	†P. florissanti	Lake Unita	1 specimen	A pine morphospecies.
incertae sedis	†Taxites	†T. eocenica	Lake Uinta	1 specimen	Isolated conifer needles of uncertain affiliation. Similar to some Cupressaceae, Pinaceae, and Taxaceae genera.

Conifer palynomorphs

Genus	Species	Member	Abundance	Notes	Image
Abies	†A. concolipites	Lake Uinta (Parachute Creek Member)	2 specimens	A true fir palynomorph similar to Abies concolor.[175]
†Abietipites	†A. antiquus		2 specimens	A generally abietine-like palyonomorph.
†Cedripites	†C. eocenicus		1 specimen	A Cedrus libani cedar-like palyonomorph.
†Inaperturopollenites	†I. concedipites		6 specimens	A Cunninghamia-like palyonomorph. First described as Cunninghamia concedipites (1933).[176]
	†I. vacuipites		1 specimen	A Chinese swamp cypress-like palyonomorph. First described as Glyptostrobus vacuipites (1933).[177]
Picea	†P. grandivescipites		6 specimens	A spruce palyonomorph.
Pinus	†P. scopulipites		Numerous	A pine palyonomorph. Less common than P. strobipites.
	†P. strobipites		Numerous	A pine palyonomorph.
	†P. tuberculipites		6 specimens	A pine palyonomorph.
Taxodium	†T. hiatipites		Numerous	A bald cypress palyonomorph.
†Tsugaepollenites	†T. viridifluminipites		1 specimen	A hemlock-like palyonomorph. First described as Tsuga viridifluminipites (1933) Moved from Zonalapollenites viridifluminipites (1953).[178] [179]

Gnetalean palynomorphs

Genus	Species	Member	Abundance	Notes	width=170px	Image
†Gnetaceae-pollenites	†G. eocenipites	Lake Uinta (Parachute Creek Member)	12 specimens	An Ephedra-like palyonomorph. First described as Ephedra eocenipites (1933).[180]

Magnoliids

Family	Genus	Species	Member	Abundance	Notes	Image
Aristolochiaceae	Aristolochia	†A. mortua			An pipevine leaf morphospecies.
Hernandiaceae	Illigera	†I. eocenica	Lake Unita		An Illigera seed morphospecies.[181]
Lauraceae	Beilschmiedia	†B. eocenica			A Beilschmiedia leaf morphospecies. Name a jr homonym of Beilschmiedia eocenica
	Lindera	†L. allardi			A spicebush leaf morphospecies.
		†L. varifolia	Lake Unita		A spicebush leaf morphospecies.
	Ocotea	†O. coloradensis	Lake Unita	2 specimens	An Ocotea leaf morphospecies. First described as Oreodaphne coloradensis (1934)
		†O. lancifolia	Lake Gosiute Lake Unita	2 specimens	An Ocotea leaf morphospecies. First described as Sapindus lancifolius (1883) Moved from Oreodaphne lancifolia (1934) Also includes Oreodaphne knowltoni (1929)
	Persea	†P. coriacea			A Persea leaf morphospecies. Name a jr homonym of Persea coriacea
Magnoliaceae	Talauma	†T. multiperforata	Lake Gosiute (Eden Valley)		An magnolia wood morphospecies.[182]

Magnoliid palynomorphs

Genus	Species	Member	Abundance	Notes	width=170px	Image
Liriodendron	†L. psilopites	Lake Uinta (Parachute Creek Member)	1 specimen	A Liriodendron tulipifera-like tulip tree palyonomorph.

Monocots

Family	Genus	Species	Member	Abundance	Notes	Image
Rowspan=8	Arecaceae	†Palmacites	†P. florissanti	Fossil Lake Lake Uinta		A palmately veined palm leaf morphospecies. Moved from Sabalites florissanti (1930) First described as Flabellaria florissanti (1885).[183]
†Palmoxylon	†P. colei	Lake Gosiute (Eden Valley)		A palm wood morphospecies.[184]
	†P. contortum	Lake Gosiute (Eden Valley)		A palm wood morphospecies.
	†P. edenense	Lake Gosiute (Eden Valley)		A palm wood morphospecies.
	†P. macginitiei	Lake Gosiute (Eden Valley)		A palm wood morphospecies.
†Phoenicites	†P. haydenii		1 specimen	A pinnately veined palm leaf morphospecies. Moved from Geonomites haydenii (1923) First described as Manicaria haydenii (1883).
Phoenix	†P. windmillis	Fossil Lake (Fossil Butte member) Lake Gosiute (Laney Member) Lake Unita (Parachute Creek Member)	numerous	A Phoenix flower morphospecies.[185]
†Sabalites	†S. powellii	Fossil Lake Lake Uinta		A palmate veined palm leaf morphospecies. First described as Sabal powellii (1883).
Cypreaceae?	†Cyperacites	†C.? filiferus	Lake Unita	A number of specimens	Likely rhizomes of monocots?. Oldest specimen first described as Diatryma? filifera, a Gastornis feather.[186] Description of the species as a bird feather was questioned by Alexander Wetmore (1930)[187] Brown (1934) moves the species to Carpolithus filiferus A second group of fossils is described as a cactus, Eopuntia douglassi (1944)[188] Brown redescribes both as cypreaceous rhizomes, and synonymized both species as Cyperus filiferus (1959)[189] Herman Becker (1963) agrees with the synonymy, but moves the species to the Cyperacites morphogenus as Cyperacites? filiferus.[190]
		†C. haydenii	Lake Gosiute	Several specimens	A sedge-like leaf morphospecies. First described as Cyperites haydenii (1885)
	Cyperus	†C. chavannesi	Rejected	1 specimen	A sedge leaf morphospecies. Rejected by Knowlton (1923) due to dubious location.
Dioscoreaceae	Dioscorea	†D. lindgrenii	Fossil Lake (Fossil Butte Member)	2 specimens	sweet potato fruits[191]
		†D. shermanii		1 specimen
Juncaceae	Juncus	Indeterminate	Rejected	?	Scrappy material tentatively identified as rush leaves. Discredited by Knowlton (1923) due to missing specimens
Poaceae	Arundo	†A. reperta	Lake Uinta	Several specimens	A giant cane grass morphospecies. The species validity of the fossils was questioned by Knowlton (1923) The species was retained by Brown (1934).
Potamogetonaceae	Potamogeton	†P. rubus	Lake Unita		A pondweed morphotype.
Typhaceae	Sparganium	†S. antiquum	Lake Gosiute Lake Uinta		A bur-reed leaf morphospecies. Moved from Brasenia? antiqua (1883) includes Pontederites hesperia (1923)
		†S. eocenicum	Lake Gosiute	1 specimen	A bur-reed leaf morphospecies.
	Typha	†T. lesquereuxi	Lake Unita		A cattail morphospecies.
Zingiberaceae	†Musophyllum	†M. complicatum	Lake Gosiute	5 specimens	A zingiberaceous monocot leaf morphotaxon.

Monocot palynomorphs

Genus	Species	Member	Abundance	Notes	width=170px	Image
†Arecipites	†A. punctatus	Lake Uinta (Parachute Creek Member)	1 specimen	A Phoenix dactylifera-like palm palyonomorph.
	†A. rugosus		1 specimen	A palm palyonomorph.
†Peltandripites	†P. davisii		1 specimen	A tuckahoe-like arrow-arum palyonomorph.
Potamogeton	†P. hollickipites		Numerous	A Potamogeton palyonomorph.
†Smilacipites	†S. echinatus		2 specimen	A Smilax-like palyonomorph.
	†S. herbaceoides		1 specimen	A Smilax-like palyonomorph.
	†S. molloides		1 specimen	A Smilax mollis-like palyonomorph.

Eudicots

Basal Eudicots

Family	Genus	Species	Member	Abundance	Notes	Image
Berberidaceae	"Mahonia"		Lake Uinta	1 Specimen	An oregon grape leaf morphospecies.[193] First described as Odostemon eocenica (1934)
Menispermaceae	†Menispermites	†M. limacioides			A moon-seed leaf morphospecies.
	Stephania	†S. wilfii	Lake Uinta (Wilkins Peak Member)	Numerous	A moon-seed fruit.[194]
Nelumbonaceae	†Nelumbo	Undescribed	Lake Gosiute		An undescribed Lotus leaf species.
Platanaceae	†Macginitiea	†M. wyomingensis			A sycamore relative species. First named Aralia macrophylla Renamed to Aralia wyomingensis (1919) Moved from Platanus wyomingensis (1969)[195]
Proteaceae	Lomatia	†L.? microphylla	Rejected	specimens missing	First identified as a guitarplant leaf morphospecies. Rejected by Brown (1934) due to missing specimens & dubious locality.
Trochodendraceae?	†Nordenskioldia	†N. borealis	Rejected		A basal trochodendroid Rejected by Manchester, Crane & Dilcher (1991) considered Carpites cocculoides like.[196]

Superasterids

Family	Genus	Species	Member	Abundance	Notes	Image
Apocynaceae	†Apocynospermum	†A. coloradensis	Lake Unita	1 specimen	An apocynaceous seed morphospecies.
	†Aspidospermoxylon	†A. uniseriatum	Lake Gosiute		An apocynaceous wood morphospecies.
Araliaceae	†Allopanax	†A. quinus	Lake Uinta (Parachute Creek Member)		An Apialean leaf mophogenus.[197] Moved from Araliophyllum quinus (1969).
Asteraceae	†Achaenites	†A. cichorioides	Lake Uinta	1 specimen	An asteraceous seed morphospecies.
Caryophyllaceae	†Alsinites	†A. revelatus	Lake Unita	1 specimen	A caryophyllacous flower morphotaxon.
Ericaceae	Andromeda	†A. delicatula		2 specimens	A possible bog rosemary ericaceous morphospecies.
Oleaceae	Osmanthus	†O. praemissa	Lake Uinta		An Osmanthus leaf morphospecies. Moved from Olea praemissa (1885)
		†O. viridiflumensis	Lake Uinta	1 specimen	An Osmanthus leaf morphospecies. Moved from Ocotea viridiflumensis (1952) First described as Oreodaphne viridiflumensis (1923)
Polemoniaceae	†Gilisenium	†G. hueberi	Fossil Lake	1 specimen	A polemoniaceous herb species.[198]
Solanaceae	†Lycianthoides	†L. calycina	Lake Uinta (Parachute Creek Member)	3 specimens	A nightshade family fruit[199]
Styracaceae	Styrax	†S. transversa	Lake Unita		A snowbell leaf morphotype.
Symplocaceae	Symplocos	†S. exilis	Lake Unita		A Symplocaceous leaf morphotype.
Incertae sedis	†Othniophyton	†O. elongatum	Lake Unita		A superastrid plant of possible caryophyllalean affinity. First described as Oreopanax elongatum (1969)[200]

Superrosids

Family	Genus	Species	Member	Abundance	Notes	Image
Altingiaceae	Liquidambar	†L. callarche	Lake Unita	2 specimens	A sweetgum species.
Anacardiaceae	†Barkleya	†B. schinoloxa	Lake Gosiute (Laney Member) Lake Uinta (Parachute Creek Member)	Numerous	An anacardiaceous samara.[201] Moved from Anacardites schinoloxus (1929). Includes Anacardites schinus Possibly the fruits of "Rhus" nigricans.
	†Edenoxylon	†E. parviareolatum	Lake Gosiute (Eden Valley)		An anacardiaceous wood morphospecies.
	Rhus	†R. longepetiolata	Lake Gosiute		An anacardiaceous leaf morphotype. Moved from Ailanthus longepetiolatus (1934)
		†"R." nigricans	Lake Gosiute (Laney Member) Lake Uinta (Parachute Creek Member)	Numerous	An anacardiaceous leaf morphotype. Moved from Myrica nigricans (1929). Possibly the leaves of Barkleya schinoloxa.
	Schmaltzia	†S. vexans	Lake Gosiute		A sumac leaf morphospecies. Moved from Rhus vexans (1885) Considered nomen dubium by MacGinitie (1969)
	†Schinoxylon	†S. actinoporosum	Lake Gosiute (Eden Valley)		An anacardiaceous wood morphospecies.
	Toxicodendron	†T. winchesteri	Lake Gosiute		A poison oak leaf morphospecies. First described as Sapindus winchesteri (1923)
Burseraceae	Bursera	†B. inaequalateralis			A bursera leaf morphotype. First named as Alnites unaequalateralis (1876)
Cannabaceae	Celtis	†C. mccoshii	Lake Gosiute Lake Unita	2 specimens	A hackberry leaf morphospecies. Includes Celtis debequensis (1928)
Capparaceae	†Forchhammerioxylon	†F. scleroticum	Lake Gosiute (Eden Valley)		A capparaceous wood morphospecies.
Celastraceae	Celastrus	†C. winchesteri	Lake Unita	1 specimen	A staff vine leaf morphospecies. Moved from Juglans winchesteri (1923) Includes Juglans alkalina (1875 [1876])
	†Celastrophyllum	†C. emarginatum	Lake Unita		A staff vine relative leaf morphospecies. MacGinitie (1969) considered the species nomen dubium.
Crassulaceae	Sedum?	†S.? hesperium	Lake Uinta	1 specimen	A possible stonecrop leaf morphospecies. Considered nomen dubium by MacGinitie (1969)
Euphorbiaceae	†Heveoxylon	†H. microporosum	Lake Gosiute (Eden Valley)		An euphorb wood morphospecies.
Fabaceae	Amorpha	†A. utensis	Lake Unita	1 specimen	A legume leaf morphospecies.
	Arcoa	†A. lindgreni	Fossil Lake	3 specimens	A caesalpinioid legume.[202]
	Caesalpinia	†C. flumen-viridensis	Lake Gosiute	2 specimens	A Caesalpinia subg. Mezoneuron seed morphospecies.[203]
		†"C." pecorae	Lake Unita	2 specimens	First named as a Caesalpinia leaf morphospecies. Considered a Detarieae tribe species by Herendeen & Dilcher (1991).
	†Caesalpinites	†C. falcatus			A Caesalpinia-like leaf morphospecies.
	Cercis	†C. tenuinervis	Lake Gosiute	1 specimen	A redbud leaf morphospecies. Considered nomen dubium by MacGinitie (1969).
	Erythrina	†E. roanensis	Lake Unita		A coral tree leaf morphospecies.
	Gymnocladus	†G. hesperia	Lake Uinta		A coffeetree leaf morphospecies. Moved from Cassia hesperia (1929) Includes Chrysobalanus lacustris (1929) & Banisteria bradleyi (1929)
	†Leguminosites	†L. alternans	Rejected		A legume leaf morphospecies. Rejected due to missing specimens by Knowlton (1923) and Brown (1934)
		†L. lesquereuxiana	Lake Unita		A legume leaf morphospecies.
		†L. regularis	Lake Unita		A legume leaf morphospecies.
	↑Parvileguminophyllum	†P. coloradensis	Lake Uinta		A legume leaf morphospecies. Moved from Mimosites coloradensis (1923).[204]
	Swartzia	†S. wardelli	Lake Uinta		A swartzieous leaf morphotype.
Fagaceae	Quercus	†Q. castaneopsis	Lake Gosiute Lake Uinta	Several fragmentary specimens	An oak morphospecies. Considered nomen dubium by MacGinitie (1969).
		†"Q. cuneatus"	Lake Unita		An oak leaf morphospecies. Name a jr homonym of Quercus cuneata
		†Q. drymeja	Rejected		An oak morphospecies. First described from Europe, Rejected from flora by MacGinitie (1969).
		†Q. petros	Lake Unita		An oak leaf morphospecies.
		†Q. utensis	Lake Unita		An oak morphospecies. Considered nomen dubium by MacGinitie (1969).
Hamamelidaceae	Distylium	†D. eocenica	Lake Unita	1 specimen	A winter-hazel leaf morphotype. Moved from Betula coloradea (1934) First named Betula eocenica (1928 non Ettingshausen (1858) Also includes Grewiopsis cissifolius (1929)
Juglandaceae	Juglans	†J. crossii	Lake Gosiute	1 specimen?	A walnut leaf morphospecies.
		†J. bilinica		1 specimen	A walnut leaf morphospecies. Moved from Hicoria juglandiformis (1898)
		†J. schimperi	Lake Unita		A walnut leaf morphospecies. Includes Juglans occidentalis (1883) & Ficus ungeri (1872)
	†Palaeocarya	Cf. †P. clarnensis	Lake Gosiute?		A Palaeocarya nut morphospecies similar to Palaeocarya clarnensis.[205]
		†P. uintaensis	Lake Unita		A Palaeocarya nut morphospecies.
	Platycarya	†P. castaneopsis	Fossil Lake Lake Uinta		A Platycarya species. Moved from Quercus castaneopsis (1883)[206]
	Pterocarya	†P. macginitii	Lake Gosiute		A wingnut seed morphospecies.[207]
		†P. roanensis	Lake Unita		A wingnut leaf morphospecies.
Malvaceae	Hibiscus	†H. roanensis	Lake Unita		An Hibiscus leaf morphospecies.
	Ochroma	†O. murata	Lake Unita	1 specimen	A balsa tree leaf morphospecies.
	Sterculia	†S. coloradensis	Lake Unita	1 specimen	A sterculioid leaf morphospecies.
	Triumfetta	T. ovata			A burbark leaf morphospecies. Jr homonym of Triumfetta ovata
Meliaceae	Cedrela	†C. trainii	Lake Unita		A meliaceous leaf morphospecies.
Moraceae	†Welkoetoxylon	†W. multiseriatum	Lake Gosiute (Tipton Shale Member)	1 specimen	A moraceous wood morphogenus.[208]
Myricaceae	Morella scalariformis	†M. scalariformis	Lake Gosiute (Eden Valley)		A myricaceous wood morphospecies. First named as Myrica scalariformis (1954) Moved from Myricoxylon scalariforme (1962)
	Myrica	†M. salicina	Rejected	sp missing	Identified as an European Myrica species Rejected by Knowlton (1923) due to the specimens being missing.
Myrtaceae	†Syzygioides	†S. americana	Lake Unita	Circa 26 specimens	A myrtaceous species. First described as Eucalyptus americana (1872) Includes Ficus omballi (1929) & Eugenia americana (1969)[209]
Rhamnaceae	†Berchemiopsis	†B. paucidentata	Lake Unita		A rhamnaceous leaf morphotype.
	Ptelea	†P. cassioides	Rejected		Type specimen leaf from the Florissant Formation Green River Fruits rejected by Call & Dilcher as not Ptelea.[210]
		†P. paliuruoides	Lake Unita		A rhamnaceous seed morphospecies. First describe as Carpolithus paliuroides (1934)
	†Rainbowia	†R. rhamnoides	Lake Unita		A rhamnaceous seed morphospecies.
Rosaceae	Prunus	†P. stewarti	Lake Gosiute		A cherry leaf morphospecies. First named Amygdalus gracilis (1883) Moved from jr homonym Prunus gracilis (non Engelm. & Gray, 1845)
	Rosa	†R. hillae			A rose leaf morphospecies.
Rutaceae	†Amyridoxylon	†A. ordinatum	Lake Gosiute (Eden Valley)		A citrus relative wood morphospecies.
	Fagara	†F. biseriata	Lake Gosiute (Eden Valley)		A citrus relative wood morphospecies.
		†F. monophylloides	Lake Gosiute (Eden Valley)		A citrus relative wood morphospecies.
Salicaceae	Populus	†P. cinnamomoides	Lake Uinta	Numerous	An Aspen/cottonwood relative. First described as Ceanothus cinnamomoides (1872) [211]
		†P. wilmattae	Lake Uinta	Numerous	An Aspen/cottonwood relative.
		†P. tidwellii	Lake Uinta	Numerous	An Aspen/cottonwood relative.
	†Pseudosalix	†P. handleyi	Lake Uinta (Parachute Creek Member)		A salicaceous species.[212] Includes Potentilla? byrami (1925), Antholithes polemonioides (1929) & Antholithes sp.6 (1934)
	Salix	†S. cockerelli	Lake Unita	4 Specimens	A willow leaf morphospecies.
		†S. longiacuminata	Lake Gosiute	1 specimen	A willow leaf morphospecies. Includes Banksia cockerelli (1929)
Sapindaceae	Allophylus	†A. flexifolia	Lake Gosiute Lake Uinta	4 Specimens	An Allophylus leaf morphospecies. First named Euonymus flexifolia (1885)
	Athyana	†A. balli	Lake Unita		A sapindaceous leaf morphotype. Moved from Rhus balli (1934).
	"Cardiospermum"		Lake Uninta	Numerous	A possible ballon vine leaf morphospecies. Moved from Lomatia coloradensis (1929)
	Koelreuteria	†K. allenii	Fossil Lake (Fossil Butte Member) Lake Unita (Parachute Creek Member)		A golden rain tree fruit morphospecies. Includes Koelreuteria uiridifluminis seeds[213]
	Sapindus	†S. dentoni	Lake Unita		A soapberry leaf morphotype. Includes Salix media (1872) & Myrica torryi (1929)
		†S. obtusifolius	Lake Unita		A soapberry leaf morphotype. Includes Salix media (1872) & Myrica torryi (1929)
	Thouinia	†T. eocenica	Lake Unita		A soap berry leaf morphospecies.
Simaroubaceae	Ailanthus	†A. lesquereuxii			A tree of heaven fruit morphospecies	upright|center|frameless
Surianaceae	Suriana	†S. inordinata	Lake Gosiute (Eden Valley)		A surianaceous wood morphospecies.
Tetramelaceae	†Parvaspicula	†P. lepidioides	Lake Uinta (Parachute Creek Member)	Numerous	A tetramelaceous fruit morphotype.[214] The type species is Clethra (?) lepidioides Cockerell (1925) Also includes Antholithes pendula (1929)
	†Punctaphyllum	†P. glandulosa	Lake Uinta (Parachute Creek Member)	Numerous	A tetramelaceous leaf morphotype. The type species is Cucurbita glandulosa (1929) Includes Aleurites glandulosa (1969) & Dendropanax latens (1974) Also includes the Ficus mississippiensis of Brown (1928)
Ulmaceae	†Cedrelospermum	†C. nervosum	Lake Uinta	Numerous	An ulmaceous species. First named as Planera nervosa (1883) [215]
Vitaceae	Cissus	†C. parrottiaefolia	Rejected	3 specimens	A grape relative leaf morphospecies. Rejected from formation by Knowlton (1923) due to distinctly different matrix.

Eudicot palynomorphs

Genus	Species	Member	Abundance	Notes	width=170px	Image
†Ailanthipites	†A. berryi	Lake Uinta (Parachute Creek Member)		An Ailanthus glandulosa-like soapberry family palynomorph.
†Alni-pollenites	†A. speciipites		Numerous	An alder-like palynomorph. first described as Alnus speciipites (1933)[216]
Betula	†B. claripites		3 specimens	A Betula lenta-like birch palynomorph.
†Caprifoliipites	†C. viridi-fluminis		1 specimen	A caprifoliaceous-like palynomorph.
Carpinus	†C. ancipites			A Carpinus-like palynomorph.
Carya	†C. juxtaporipites		1 specimen	A hickory-like palynomorph. First described as Hicoria juxtaporipites (1933)[217]
	†C. viridi-fluminipitis		1 specimen	A Carya myristiciformis-like hickory palynomorph. First described as Hicoria viridi-fluminipites (1933)[218]
Ericipites	†E. brevisulcatus		1 specimen	A Calluna vulgaris-like heath family palynomorph.
	†E. longisulcatus		2 specimens	A heath family-like palynomorph.
†Juglanspollenites	†J. nigripites		1 specimen	A walnut-like palynomorph. First described as Juglans nigripites (1933)[219]
†Momipites	†M. corylipites		2 specimens	An Engelhardia-like palynomorph. First described as Engelhardia corylipites (1933).[220]
	†M. coryloides		Numerous	A Celtis iguanaea-like hackberry palynomorph.
†Myricipites	†M. dubius		1 specimen	A bayberry-like palynomorph.
Myriophyllum	†M. ambiguipites		1 specimen	A Myriophyllum spicatum-like watermillfoil family palynomorph.
†Rhoipites	†R. bradleyi		4 specimens	A Rhus typhina-like cashew family palynomorph.
Salix	†S. discoloripites		Numerous	A Salix discolor and S. fragilis-like willow palynomorph.
†Talisiipites	†T. fischeri		Numerous	A Talisia depressa-like soapberry family palynomorph.
Tilia	†T. crassipites		1 specimen	A Tilia americana-like linden palynomorph.
	†T. tetraforaminipites		3 specimens	A Tilia americana-like linden palynomorph.
	†T. vescipites		1 specimens	A Tilia americana-like linden palynomorph.
†Vitipites	†V. dubius		1 specimen	A Vitis vinifera-like grape family palynomorph.

Angiosperms of uncertain affiliation

Genus	Species	Member	Abundance	Notes	width=170px	Image
†Achaenites	†A. cichorioides			A achene like structure of unidentified affiliation.
†Antholithes	†A. dubia	Lake Unita	1 specimen	An 8-lobed/petaled flower of unidentified affiliation.
	†A. improbus		2 specimens	A flowering structure of unidentified affiliation.
	†A. vitaciflora	Lake Unita	1 specimen	A cluster of possible vitaceous flowers-fruits.
	†A. winchesteri	Lake Unita	1 specimen	A flowering structure of unidentified affiliation. First described as Sambucus? winchesteri (1923)
	†A. sp 1	Fossil Lake	1 specimen	A possibly cruciferous or orrograceous flower of unidentified affiliation.
	†A. sp 2	Lake Unita	1 specimen	A flower with many stamens of unidentified affiliation.
	†A. sp 3	Lake Unita	1 specimen	A malvaceous looking flower of unidentified affiliation.
	†A. sp 4	Lake Unita	1 specimen	A flower with a superior ovary surrounded by the five-toothed calyx.
	†A. sp 5	Lake Unita	1 specimen	A flower with tubular receptacle surmounted by a four-parted calyx.
	†A. sp 7	Fossil Lake	1 specimen	A large flower or a badly distorted leaf of a Nelumbo?
†Bonanzacarpum	†B. sprungerorum	Lake Uinta (Parachute Creek Member)	Numerous	A eudicot fruit of uncertain phylogenetic placement.[221]
†Carpites	†C. inquirenda	Lake Unita	1 specimen	A nutlet or fruit of unidentified affiliation.
	†C. newberryanus		3 specimens	A nutlet of unidentified affiliation. First described as Juglans occidentalis nuts (1898). First spelled Carpites newberryana
	†C. viburni		1 specimen	A seed or nutlet of unidentified affiliation. Includes Carpolithes capsularis (1929)
†Carpolithus	†C. alabaccus	Lake Unita	1 specimen	A winged or membranous fruit of unidentified affiliation.
	†C. caryophylloides	Lake Unita		A dry capsular fruit of unidentified affiliation.
	†C. cassioides	Lake Unita	2 specimens?	A legume like pod of unidentified affiliation.
	†C. insectoides	Lake Unita	1 specimen	A winged fruit of unidentified affiliation.
	†C. palrnites	Lake Unita	1 specimen	A palm fruit or husk like fossil of unidentified affiliation.
	†C. prunoides	Lake Unita	1 specimen	A flattened fruit of unidentified affiliation. First named Carpolithus ellipticus
	†C. serratifolius	Lake Unita	1 specimen	Branchlet with leaves and fruits of possible salicaceous affinity.
	†C. vitaceus	Lake Unita	1 specimen	A possibly vitaceous seed of unidentified affiliation.
†Caulinites	†C. acanthus	Lake Unita	1 specimen	A thorny stem of unidentified affiliation.
	†C. prehensus	Lake Unita	1 specimen	A climbing plant tendril of unidentified affiliation.
†Chaneya	†C. tenuis	Lake Gosiute Lake Unita		A sapindalean flower of uncertain affiliations. Includes Astronium truncatum (1953).[222]
†Dicotylophyllum	†D. lesquereuxii		1 specimen	A leaf of uncertain affiliation. First described as Acer indivisum Moved from Acer lesquereuxii [223]
†Palibinia	†P. comptonifolia	Lake Uinta (Parachute Creek Member)		A pentapetalean eudicot of uncertain affiliation.[224] Moved from Vauquelinia comptonifolia (1969) Originally named Banksia comptonifolia (1934)
†Phyllites	†P. ericoides	Lake Unita	1 specimen	A thick leaf of unidentified affiliation.
	†P. fremonti	Rejected	1 specimen	A leaf of unidentified affiliation. R.W. Brown (1934) rejected due to doubtful locality and lost specimen.
	†P. glandiferus	Lake Unita	1 specimen	An odd-pinnately compound leaf of possible fabaceous affinity.
	†P. juncoides	Lake Unita	1 specimen	A leaf of unidentified affiliation.
	†P. ouratea	Lake Unita	1 specimen	A possibly herbaceous leaf of unidentified affinity.
	†P. winchesteri	Lake Unita	1 specimen	A leaf of unidentified affiliation.

Plants of uncertain affiliation

Genus	Species	Member	Abundance	Notes	width=170px	Image
†Rhizocaulon	†R. dichotomum	Lake Unita	1 specimen	A root of unidentified affiliation. Has a distinct branched fork.
	†R. natans	Lake Unita	1 specimen	A rootlet of unidentified affiliation. small with Trapa natans-like rhizoids.

Notes and References

1. De Carvalho . Marcelo R. . Maisey . John G. . Grande . Lance . June 2004 . Freshwater Stingrays of the Green River Formation of Wyoming (Early Eocene), With the Description of a New Genus and Species and an Analysis of Its Phylogenetic Relationships (Chondrichthyes: Myliobatiformes) . Bulletin of the American Museum of Natural History . en . 284 . 1–136 . 10.1206/0003-0090(2004)284<0001:FSOTGR>2.0.CO;2 . 0003-0090.
2. Book: Grande . L. . 2013 . The lost world of fossil lake: snapshots from deep time . University of Chicago Press; Illustrated edition . 9780226922966 .
3. Schwartz . Frank J. . 2011-10-01 . An Emperical Synthetic Pattern Study of Gars (Lepisosteiformes) and Closely Related Species, Based Mostly on Skeletal Anatomy the Resurrection of Holostei . Journal of North Carolina Academy of Science . en . 127 . 3 . 219 . 10.7572/2167-5880-127.3.219 . 2167-5880.
4. Grande . L. . 1982 . A revision of the fossil genus Knightia, with a description of a new genus from the Green River Formation (Teleostei, Clupeidae) . American Museum Novitates . 2731 . 1–22 .
5. Near . Thomas J. . Thacker . Christine E. . 2024-04-18 . Phylogenetic Classification of Living and Fossil Ray-Finned Fishes (Actinopterygii) . Bulletin of the Peabody Museum of Natural History . 65 . 1 . 10.3374/014.065.0101 . 0079-032X. free .
6. Web site: Fossil Fish - Fossil Butte National Monument (U.S. National Park Service) . 2024-08-17 . www.nps.gov . en.
7. 10.1371/journal.pone.0010420. free. First Known Feeding Trace of the Eocene Bottom-Dwelling Fish Notogoneus osculus and Its Paleontological Significance. 2010. Martin. Anthony J.. Vazquez-Prokopec. Gonzalo M.. Page. Michael. PLOS ONE. 5. 5. e10420. 20463969. 2864752. 2010PLoSO...510420M.
8. Henrici . Amy C. . Báez . Ana M. . Grande . Lance . 2013 . Aerugoamnis paulus, New Genus and New Species (Anura: Anomocoela): First Reported Anuran from the Early Eocene (Wasatchian) Fossil Butte Member of the Green River Formation, Wyoming . Annals of Carnegie Museum . en . 81 . 4 . 295–309 . 10.2992/007.081.0402 . 0097-4463.
9. Rieppel . O. . Grande . L. . 1998 . A well-preserved fossil amphiumid (Lissamphibia: Caudata) from the Eocene Green River Formation of Wyoming . Journal of Vertebrate Paleontology . 18 . 4 . 700–708 . 10.1080/02724634.1998.10011099. 1998JVPal..18..700R .
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11. Web site: PBDB Taxon . 2024-08-17 . paleobiodb.org.
12. Carpenter . K. . 2018-10-18 . Soft-bodied fossil of a lizard from the Parachute Creek Member, Green River Formation (Eocene), Utah . Geology of the Intermountain West . en . 5 . 263–269 . 10.31711/giw.v5.pp263-269 . 2380-7601. free .
13. Conrad . J.L. . Head . J.J. . Carrano . M.T. . 2014 . Unusual soft-tissue preservation of a crocodile lizard (Squamata, Shinisauria) from the Green River Formation (Eocene) and shinisaur relationships . The Anatomical Record . 297 . 3 . 545–559. 10.1002/ar.22868 . 24482393 .
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15. Hekkala . E. . Gatesy . J. . Narechania . A. . Meredith . R. . Russello . M. . Aardema . M. L. . Jensen . E. . Montanari . S. . Brochu . C. . Norell . M. . Amato . G. . 2021-04-27 . Paleogenomics illuminates the evolutionary history of the extinct Holocene "horned" crocodile of Madagascar, Voay robustus . Communications Biology . en . 4 . 1 . 505 . 10.1038/s42003-021-02017-0 . 33907305 . 8079395 . 2399-3642 . free.
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