List of non-marine molluscs of Vietnam explained

The non-marine molluscs of Vietnam are a part of the molluscan fauna of Vietnam (wildlife of Vietnam). A number of species of non-marine mollusks are found in the wild in Vietnam.

There are good reasons to suppose that Vietnam, with a surface of 332,000 km², a large variety of habitats, and many different limestone 'islands' that differ from each other in faunal composition,^[1] will have a rich diversity of terrestrial molluscs. Numerous non-marine mollusc species, including more than 850 species of land gastropods,^[2] have been described from the country but many others still await discovery and description.

Freshwater gastropods

Freshwater gastropods in Vietnam include:

Neritidae

• Neritina violacea

Ampullariidae

• Pila polita
• Pila conica
• Pila ampullacea

• Pomacea canaliculata
• Pomacea insularum (d'Orbigny, 1835)

Viviparidae

• Cipangopaludina lecythoides^[3]
• Idiopoma umblicata^[3]
• Sinotaia aeruginosa^[3]
• Mekongia lithophaga^[3]
• Mekongia hainesiana^[3]
• Filopaludina sumatrensis^[3]
• Angulyagra polyzonata^{[4] [3]}
• Angulyagra duchieri^[3]
• Angulyagra boettgeri^[3]

Pachychilidae

• Adamietta housei^[3]
• Adamietta reevei^[3]
• Brotia annamita^[3]
• Brotia dautzenbergiana^[3]
• Brotia hoabinhensis^[3]
• Brotia swinhoei^[3]
• Sulcospira collyra^[3]
• Sulcospira dakrongensis^[3]
• Sulcospira hainanensis^[3]
• Sulcospira proteus^[3]
• Sulcospira quangtriensis^[3]
• Sulcospira tonkiniana^[3]
• Sulcospira touranensis^[3]
• Sulcospira vietnamensis^[3]
• Semisulcospira aubryana^[3]
• Paracrostoma paludiformis^[3]

Paludomidae

• Paludomus messageri (Bavay & Dautzenberg, 1900)^[3]

Potamididae (brackish water snails)

• Cerithidea sp.^[3]

Thiaridae

• Melanoides tuberculata (O. F. Müller, 1774)^{[5] [4] [3]}
• Neoradina prasongi^[3]
• Sermyla requetii^[3] - synonym: Sermyla tornatela^{[4] [3]}
• Tarebia granifera (Lamarck, 1822)^{[4] [3]}
• Thiara scabra^{[4] [3]}

Assimineidae

• Assiminea brevicula^[3]
• Assiminea francoisi^[3]
• Assiminea interrupta^[3]
• Assiminea obtusa^[3]
• Assiminea lutea^[3]

Bithyniidae

• Digoniostoma siamensis^[3]
• Parafossarulus striatulus - synonym: Parafossarulus manchouricus^[3]
• Alocinma longicornis^[3]
• Bithynia misella^[3]
• Bithynia fuchsiana^{[4] [3]}
• Bithynia goniomphalos^[3]
• Wattebledia crosseana^[3]

Pomatiopsidae

• Neoprososthenia levayi - synonym: Paraprososthenia levayi^[3]
• Lacunopsis harmandi^[3]
• Tricula ovata^[3]
• Tricula similunaris^[3]
• Vietricula - 11 species^[3]

Stenothyridae

• Stenothyra divalis^[3]
• Stenothyra messageri^{[6] [4] [3]}
• Stenothyra monilifera^[3]
• Stenothyra polita^[3]
• Stenothyra cyrtochila^[3]
• Stenothyra ovata^[3]
• Stenothyra conica^[3]
• Stenothyra alba^[3]
• Stenothyra schlickumi Brandt, 1968^[3]

Iravadiidae (marine/brackish)

• Fluvicingula elegantula (A. Adams, 1861)^[3]
• Iravadia cochinchinensis (Bavay & Dautzenberg, 1910)^[3]
• Iravadia ornata Blanford, 1867^[3]
• Iravadia quadrasi (O. Boettger, 1893)^[3]
• Iravadia rohdei (Brandt, 1968)^[3]
• Pseudomerelina mahimensis (Melvill, 1893)^[3]
• Hyala sp.^[3]

Buccinidae

• Clea helena (Meder in Philippi, 1847)^[3]

Lymnaeidae

• Austropeplea viridis (Quoy & Gaimard, 1832)^[7]
• Radix auricularia (Linnaeus, 1758) - synonym: Lymnaea swinhoei^[7]
• Radix rubiginosa (Michelin, 1831)^[7]

Planorbidae

• Polypylis hemisphaerula (Benson, 1842)^[4]
• Gyraulus convexiusculus (Hutton, 1849)^[4]

Land gastropods

Land gastropods in Vietnam include:

Cyclophoridae

• Chamalycaeus depressus (Bavay et Dautzenberg, 1912)^[8]
• Chamalycaeus fraterculus (Bavay et Dautzenberg, 1900)^[8]
• Chamalycaeus heudei (Bavay et Dautzenberg, 1900)^[8]
• Chamalycaeus paviei (Bavay et Dautzenberg, 1912)^[8]
• Dioryx compactus (Bavay et Dautzenberg, 1900)^[8]
• Dioryx messageri (Bavay et Dautzenberg, 1900)^[8]
• Dioryx vanbuensis (Bavay et Dautzenberg, 1900)^[8]
• Laotia christahemmenae Páll-Gergely, 2014^[8]
• Cyclophorus affinis Theobald, 1858^[9]
• Cyclophorus amoenus (Pfeiffer, 1854)^[9]
• Cyclophorus aquilus (Sowerby, 1843)^[9]
• Cyclophorus aurantiacus pernobilis Gould, 1844^[9]
• Cyclophorus bensoni (Pfeiffer, 1854)^[9]
• Cyclophorus cambodgensis Morlet, 1884^{[9] [8]}
• Cyclophorus cantori (Benson, 1851)^[9]
• Cyclophorus clouthianus Möllendorff, 1882^[9]
• Cyclophorus consociatus Smith, 1893^[9]
• Cyclophorus courbeti Ancey, 1888^[10]
• Cyclophorus courbeti courbeti Ancey, 1888^[9]
• Cyclophorus courbeti leucostoma Dautzenberg et Fischer, 1905^[9]
• Cyclophorus cucphuongensis Oheimb, 2019^[10]
• Cyclophorus cryptomphalus Benson, 1857^[9]
• Cyclophorus dilatatus Heude, 1886^[9]
• Cyclophorus dodrans dodrans Mabille, 1887^[9]
• Cyclophorus dodrans fasciatus Kobelt, 1908^[9]
• Cyclophorus donghoiensis Thach et Huber, 2017^[9]
• Cyclophorus eudeli Smith, 1893^[9]
• Cyclophorus expansus (Pfeiffer, 1853)^[9]
• Cyclophorus fargesianus Heude, 1885^[9]
• Cyclophorus fasciatus Kobelt, 1908^[10]
• Cyclophorus floridus (Pfeiffer, 1854)^[9]
• Cyclophorus fruhstorferi fruhstorferi Möllendorff, 1901^[9]
• Cyclophorus fruhstorferi langsonensis Kobelt, 1908^[9]
• Cyclophorus fulguratus Pfeiffer, 1852^[9]
• Cyclophorus haughtoni Theobald, 1858^[9]
• Cyclophorus herklotsi Martens, 1861^[9]
• Cyclophorus huberi Thach, 2016^[9]
• Cyclophorus ignilabris Möllendorff, 1901^[9]
• Cyclophorus implicatus Bavay et Dautzenberg, 1908^[8]
• Cyclophorus implicatus implicatus Bavay et Dautzenberg, 1908^[9]
• Cyclophorus implicatus kanhoensis Do et Do, 2019^[9]
• Cyclophorus jourdyi Morlet, 1886^{[10] [9]}
• Cyclophorus labiosus (Pfeiffer, 1854)^[9]
• Cyclophorus malayanus (Benson, 1852)^[9]
• Cyclophorus mansuyi Dautzenberg et Fischer, 1908^[9]
• Cyclophorus martensianus Möllendorff, 1874^[9]
• Cyclophorus massiei Morlet, 1891^{[9] [8]}
• Cyclophorus monachus (Morelet, 1866)^[9]
• Cyclophorus muspratti Godwin-Austen et Beddome, 1894^[9]
• Cyclophorus orthostylus Möllendorff, 1898^[9]
• Cyclophorus paracucphuongensis Oheimb, 2019^[10]
• Cyclophorus paviei Morlet, 1884^[9]
• Cyclophorus pfeifferi Reeve, 1861^[9]
• Cyclophorus phongnhakebangensis Oheimb, 2019^[10]
• Cyclophorus polystictus Möllendorff, 1901^[9]
• Cyclophorus pyrostoma Möllendorff, 1882^[9]
• Cyclophorus saturnus Pfeiffer, 1862^[9]
• Cyclophorus siamensis (Sowerby, 1850)^[9]
• Cyclophorus songmaensis Morlet, 1891^[9]
• Cyclophorus speciosus (Philippi, 1847)^[9]
• Cyclophorus subfloridus Ancey, 1888^{[10] [9]}
• Cyclophorus takumisaitoi Hirano, 2019^[10]
• Cyclophorus tamdaoensis Do et Do, 2019^[9]
• Cyclophorus tetrachrous Mabille, 1887^[9]
• Cyclophorus theodori Ancey, 1888^[9]
• Cyclophorus tornatus Morlet, 1892^[9]
• Cyclophorus trouiensis trouiensis Wattebled, 1886^[9]
• Cyclophorus trouiensis bendei Varga, 1972^[9]
• Cyclophorus trouiensis omphalotropis Möllendorff, 1900^[9]
• Cyclophorus unicus Mabille, 1887^[9]
• Cyclophorus volvulus (O.F. Müller, 1774)^{[9] [8]}
• Cyclophorus zebrinus (Benson, 1836)^[9]
• Cyclotus lubricus (Dautzenberg et Fischer, 1908)^[8]
• Japonia fischeri (Morlet, 1886)^[8]
• Japonia scissimargo (Benson, 1856)^[8]
• Platyrhaphe leucacme Möllendorff, 1901^[8]
• Platyraphe sordida (Pfeiffer, 1855)^[8]
• Pterocyclos prestoni Bavay et Dautzenberg, 1909^[8]
• Rhiostoma morleti Dautzenberg et Fischer, 1905^[8]
• Scabrina laciniana (Heude, 1885)^[8]
• Scabrina locardi (J. Mabille, 1887)
• Scabrina tonkiniana (J. Mabille, 1887)^[11]
• Scabrina vanbuensis (Smith, 1896)^{[8] [11]}

Diplommatinidae

• Diplommatina balansai Morlet, 1886^[8]
• Diplommatina clausilioides Bavay et Dautzenberg, 1912^[8]
• Diplommatina demangei Bavay et Dautzenberg, 1912^[8]
• Diplommatina messageri Ancey, 1903^[8]

Pupinidae

• Pupina anceyi Bavay & Dautzenberg, 1899^{[12] [8]}
• Pupina billeti Fischer, 1898^[12]
• Pupina brachysoma Ancey, 1903^[12]
• Pupina dorri Dautzenberg, 1893^{[12] [8]}
• Pupina douvillei Dautzenberg & Fischer, 1905^[12]
• Pupina exclamationis Mabille, 1887^{[12] [8]}
• Pupina flava Möllendorff, 1884^[12]
• Pupina illustris Mabille, 1887^[12]
• Pupina laffonti Ancey, 1899^[12]
• Pupina solidula Möllendorff, 1901^[12]
• Pupina sonlaensis Do, 2017^[12]
• Pupina thaitranbaii Do, 2017^[12]
• Pupina tonkiana Bavay & Dautzenberg, 1899^{[12] [8]}
• Pupina verneaui Dautzenberg & Fischer, 1905^{[12] [8]}
• Pupina vescoi Morelet, 1862^[12]
• Pseudopomatias amoenus Möllendorff, 1885^[8]
• Pseudopomatias maasseni Páll-Gergely et Hunyadi, 2015^[8]
• Pseudopomatias sophiae Páll-Gergely, 2015^[8]
• Pupinella mansuyi (Dautzenberg et Fischer, 1908)^[8]
• Rhaphaulus tonkinensis Páll-Gergely, 2014^[8]
• Streptaulus longituba Páll-Gergely & Gargominy, 2017^[13]

Helicinidae

• Geotrochatella jourdyi Dautzenberg, 1895^[8]

Hydrocenidae

• Georissa decora Möllendorff, 1900^[8]

Clausiliidae

• Castanophaedusa fontainei Páll-Gergely & Szekeres, 2017^[14]
• Castanophaedusa huberi (Thach, 2016)^[14]
• Garnieria mouhoti nhuongi Do, 2015^[14]
• Grandinenia ardouiniana (Heude, 1885)^[14]
• Grandinenia tonkinensis (Nordsieck, 2010)^[14]
• Hemiphaedusa babeensis crassitesta Nordsieck, 2011^[15]
• Hemiphaedusa porphyrostoma regina Nordsieck, 2011^[15]
• Hemiphaedusa thatkheana ptychostoma Nordsieck, 2011^[15]
• Hemiphaedusa thatkheana splendida Nordsieck, 2011^[15]
• Leptacme cuongi Maassen & Gittenberger, 2007^[16]
• Leptocochlea sykesi lethrungtongi Grego & Szekeres, 2011^[17]
• Liparophaedusa leptotesta Nordsieck, 2011^[15]
• Megalauchenia soror Nordsieck, 2011^[15]
• Messageriella gargominyi Páll-Gergely & Szekeres, 2017^[14]
• Oospira abstrusa Szekeres, 1970^[14]
• Oospira abstrusa ginkae Grego & Szekeres, 2014^[14]
• Oospira duci Maassen & Gittenberger, 2007^[16]
• Oospira duci khanhi Nordsieck, 2011^[15]
• Oospira haivanensis Chinh & Szekeres, 2019^[18]
• Oospira naggsi Luong & Szekeres, 2014^[14]
• Oospira naggsi naggsi Luong & Szekeres, 2014^[14]
• Oospira naggsi parva Páll-Gergely & Szekeres, 2017^[14]
• Oospira oviformis Nordsieck, 2011^[15]
• Oospira pyknosoma Gittenberger & Vermeulen, 2001^[19]
• Oospira smithi Maassen & Gittenberger, 2007^[16]
• Oospira triptyx Nordsieck, 2011^[15]
• Phaedusa cochinchinensis (Pfeiffer, 1841)^[18]
• Phaedusa lypra pereupleura Nordsieck, 2011^[15]
• Phaedusa micropaviei Nordsieck, 2011^[15]
• Synprosphyma oospiroides Nordsieck, 2011^[15]
• Tropidauchenia bavayi senescens Nordsieck, 2011^[15]
• Tropidauchenia (Euryauchenia) fisheri phasmoides Grego & Szekeres, 2011^[17]
• Tropidauchenia ootanii longicollis Nordsieck, 2011^[15]
• Tropidauchenia palatalis Nordsieck, 2011^[15]

Helicarionidae

• Falsiplecta integripedia Schileyko & Semenyuk, 2018^[20]
• Laocaia attenuata Kuzminykh, 1999^[21]
• Laocaia obesa Kuzminykh, 1999^[21]
• Laocaia simovi Dedov & Schneppat, 2019^[21]

Streptaxidae - 39 species of Streptaxidae are known from Vietnam^[22]

• Discartemon^[22]
• Indoartemon^[22]
• Perrottetia aberrata (Souleyet, 1852)^[22]
• Perrottetia cristatellus (Möllendorff, 1901)^[22]
• Perrottetia daedaleus (Bavay & Dautzenberg, 1908)^[22]
• Perrottetia dugasti (Morlet, 1892)^[22]
• Perrottetia gudei (Fulton, 1915)^[22]
• Perrottetia hongthinhae D.-S. Do, 2017^[22]
• Perrottetia mabillei (Bavay et Dautzenberg, 1903)^[22]
• Perrottetia messageri (Bavay et Dautzenberg, 1908)^[22]
• Perrottetia oppidulum (Bavay et Dautzenberg, 1908)^[22]
• Perrottetia simonianus (Heude, 1890)^[22]
• Stemmatopsis arcuatolabris Do, 2021^[23]
• Stemmatopsis nangphaiensis Do & Do, 2015^[23]
• Stemmatopsis vanhoensis Do & Do, 2015^[23]
• Haploptychius^[22]
• Oophana^[22]
• Elma Adams, 1866^{[24] [22]}
• Huttonella^[22]

Plectopylidae - 16 species are known from Vietnam

• Gudeodiscus anceyi (Gude, 1901)
• Gudeodiscus cyrtochilus (Gude, 1909)
• Gudeodiscus dautzenbergi (Gude, 1901)
• Gudeodiscus fischeri (Gude, 1901)
• Gudeodiscus francoisi (Fischer, 1898)
• Gudeodiscus giardi (Fischer, 1898)
• Gudeodiscus hemmeni Páll-Gergely & Hunyadi, 2015^[25]
• Gudeodiscus infralevis (Gude, 1908)^[25]
• Gudeodiscus messageri (Gude, 1909)^[25]
• Gudeodiscus phlyarius (Mabille, 1887)^[25]
• Gudeodiscus suprafilaris (Gude, 1908)^[25]
• Gudeodiscus villedaryi (Ancey, 1888)^[25]
• Gudeodiscus emigrans (Möllendorff, 1901)^[25]
• Halongella fruhstorferi (Möllendorff, 1901)^[25]
• Halongella schlumbergeri (Morlet, 1886)^[25]
• Sicradiscus mansuyi (Gude, 1908)^[25]

Philomycidae

• Meghimatium bilineatum (Benson, 1842)^[26]

Ariophantidae

• Hemiplecta jensi Páll-Gergely, 2019^[27]
• Parmarion martensi Simroth, 1894^[26]
• Parmarion pupillaris Humbert, 1864^[26]

Agriolimacidae

• Deroceras laeve (O. F. Müller, 1774)^[28]

Camaenidae

• Amphidromus cambojiensis (Reeve, 1860)^[29]
• Amphidromus ingens Möllendorff, 1900^[29]
• Amphidromus naggsi Thach & Huber, 2014^[29]

Freshwater bivalves

Freshwater bivalves in Vietnam include:

See also

• List of marine molluscs of Vietnam

Lists of molluscs of surrounding countries:

• List of non-marine molluscs of China
• List of non-marine molluscs of Laos
• List of non-marine molluscs of Cambodia

References

This article incorporates CC-BY-3.0 text from the reference^[16]

Further reading

• Do D. S., Nguyen T. H. T. & Do V. N. (2015). "A checklist and classification of terrestrial prosobranch snails from Son La, north-western Vietnam". Ruthenica 25(4): 117–132. PDF.
• Vermeulen J. J. & Whitten T. (1998). "Land and freshwater molluscs of the karst regions ENE of Haiphong and the Cuc Phuong National Park, northern Vietnam". Report of a survey for the Vietnam Programme of FFI 12 pp. (unpublished report).
• Vermeulen J. J. & Maassen W. J. M. (2003). "The non-marine mollusk fauna of the Pu Luong, Cuc Phuong, Phu Ly, and Ha Long regions in northern Vietnam". Report of a survey for the Vietnam Programme of FFI 35 pp. (unpublished report).

Notes and References

1. Oheimb . Parm Viktor von . Oheimb . Katharina C. M. von . Hirano . Takahiro . Do . Tu Van . Luong . Hao Van . Ablett . Jonathan . Pham . Sang Van . Naggs . Fred . 2018 . Competition matters: Determining the drivers of land snail community assembly among limestone karst areas in northern Vietnam . . 8 . 8. 4136–4149 . 29721286 . 10.1002/ece3.3984 . 5916308 .
2. Book: Raheem . Dinarzarde C. . Backeljau . Thierry . Pearce-Kelly . Paul . Taylor . Harry . Fenn . Jonathan . Sutcharit . Chirasak . Panha . Somsak . von Oheimb . Katharina C. M. . von Oheimb . Parm Viktor . Ikebe . Chiho . Páll-Gergely . Barna . Gargominy . Olivier . Lương . Hào Văn . Phạm . Sáng Văn . Đỗ . Tứ Văn . Đinh . Phòng Thị . Naggs . Manel . Ablett . Jon . Mackenzie Dodds . Jackie . Wade . Christopher M. . Naggs . Fred . 2017 . An Illustrated Guide to the Land Snails and Slugs of Vietnam . The Natural History Museum, the Royal Belgian Institute of Natural Sciences & the Zoological Society of London . 978-0-565-09458-4.
3. Madsen . H. . Hung . N. M. . 2015 . Reprint of "An overview of freshwater snails in Asia with main focus on Vietnam" . . 141 . Pt B . 372–384 . 10.1016/j.actatropica.2014.10.014 . 25446169 .
4. Dung . B. T. . Madsen . H. . The . D. T. . 2010 . Distribution of freshwater snails in family-based VAC ponds and associated waterbodies with special reference to intermediate hosts of fish-borne zoonotic trematodes in Nam Dinh Province, Vietnam . . 116 . 1. 15–23 . 10.1016/j.actatropica.2010.04.016 . 20457118 .
5. Madhyastha A. (2010). Melanoides tuberculatus. In: IUCN 2010. IUCN Red List of Threatened Species. Version 2010.4. . Downloaded on 3 December 2010.
6. Phan . V. T. . Ersbøll . A. K. . Nguyen . K. V. . Madsen . H. . Dalsgaard . A. . 2010 . Farm-Level Risk Factors for Fish-Borne Zoonotic Trematode Infection in Integrated Small-Scale Fish Farms in Northern Vietnam . . 4 . 7. e742 . 10.1371/journal.pntd.0000742 . 20644617 . 2903470 . free .
7. Dung . B. T. . Doanh . P. N. . The . D. T. . Loan . H. T. . Losson . B. . Caron . Y. . 2013 . Morphological and Molecular Characterization of Lymnaeid Snails and Their Potential Role in Transmission of Fasciola spp. in Vietnam . . 51 . 6. 657–662 . 10.3347/kjp.2013.51.6.657 . 24516270 . 3916454 .
8. DO Duc Sang, NGUYEN Thi Hong Thinh & DO Van Nhuong (2015). "A checklist and classification of terrestrial prosobranch snailsfrom Son La, north-western Vietnam". Ruthenica 25(4): 117-132.
9. DO Duc Sang, DO Van Nhuong (2019). "Family Cyclophoridae in Vietnam (Gastropoda: Cyclophoroidea): the genus Cyclophorus Montfort, 1810". Ruthenica 29(1): 1-53.
10. Oheimb . Katharina C. M. von . Oheimb . Parm Viktor von . Hirano . Takahiro . Do . Tu Van . Ablett . Jonathan . Luong . Hao Van . Pham . Sang Van . Naggs . Fred . 2019 . Cryptic diversity of limestone karst inhabiting land snails (Cyclophorus spp.) in northern Vietnam, their evolutionary history and the description of four new species . . 14 . 10. e0222163 . 10.1371/journal.pone.0222163 . 31644567 . 6808330 . 2019PLoSO..1422163V . free .
11. Junn Kitt Foon & Mohammad Effendi Marzuki (2022). "First record and description of a new Scabrina species (Gastropoda: Cyclophoridae) from Peninsular Malaysia". Folia Malacologica 1(30): 23-35. DOI: https://doi.org/10.12657/folmal.030.003
12. Do. D.S.. Two new species of the genus Pupina (Caenogastropoda: Pupinidae) from Northwestern Vietnam. Raffles Bulletin of Zoology. 2017. 65. 299–303.
13. Barna Páll-Gergely, Olivier Gargominy, Benoît Fontaine & Takahiro Asami (2017). "Breathing device of a new Streptaulus species from Vietnam extends understanding of the function and structure of respiratory tubes in cyclophoroids (Gastropoda: Caenogastropoda: Pupinidae)". Journal of Molluscan Studies 83(2): 243-248. doi:10.1093/mollus/eyx006
14. Barna Páll-Gergely & Miklós Szekeres (2017). "New and little-known Clausiliidae (Gastropoda: Pulmonata) from Laos and Southern Vietnam". Journal of Conchology 42(6): 507–521.
15. Hartmut Nordsieck (2011). Clausiliidae of Vietnam with the description of new taxa (Gastropoda: Stylommatophora. Archiv für Molluskenkunde 140(2): 149-173. DOI:10.1127/arch.moll/1869-0963/140/149-173
16. Maassen W. J. M. & Gittenberger E. (2007). "Three new clausiliid land snails from Tonkin, northern Vietnam (Gastropoda: Pulmonata: Clausiliidae)". Zoologische Mededelingen 81(1): http://www.zoologischemededelingen.nl/81/nr01/a10
17. Jozef Grego & Miklos Szekeres (2011). "New Taxa of Asiatic Clausiliidae (Mollusca: Gastropoda)". Visaya 3(2): 4-22.
18. Bui Thi Chinh & Miklós Szekeres (2019). "A new species of the genus Oospira Blanford, 1872 (Gastropoda, Pulmonata, Clausiliidae) from central Vietnam". Ruthenica 29(4): 185-189.
19. Gittenberger . E. . Vermeulen . J. J. . 2001 . Oospira (O.) pyknosoma spec. nov. (Gastropoda, Pulmonata, Clausiliidae), an impressive clausiliid species from Vietnam . . 65 . 123–129 .
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21. Ivaylo Dedov, Ulrich Schneppat, Manh Quang Vu & Nguyen Quoc Huy (2019). "A new semislug of the genus Laocaia (Gastropoda, Pulmonata, Helicarionidae) from Vietnam". ZooKeys 846: 19-30.
22. 10.12657/folmal.025.009. 25. 2. 95–100. Sang. Do Duc. Do Duc. Sang. Checklist of the genus Perrottetia Kobelt, 1905 (Pulmonata: Streptaxidae) of Vietnam, with description of a new species. Folia Malacologica. 2017-11-02. 2017. free.
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