Calcare di Aurisina explained

Calcari di Aurisina
Type:Geological formation
Age:Santonian-Campanian
~
Period:Campanian
Prilithology:Limestone
Namedfor:Aurisina
Region:Trieste
Coordinates:45.8°N 13.6°W
Paleocoordinates:23.5°N 16.2°W
Subunits:Zolla member
Area:Friuli-Venezia Giulia

The Calcare di Aurisina (Also known as Aurisina Formation, “Formazione di Monrupino”, Trieste Karst Limestone formation, “Repen formation”, “Sežana formation” and “Lipiza formation”)[1] [2] [3] is a Mesozoic geologic formation in Italy and Slovenia. This limestones are found in the Trieste area and are of Late Cretaceous (Santonian-Campanian) age, being a local record of Carbonate platform limestones, historically quarried by cutting large blocks using steel wire, what has allowed more access to them, as numerous quarries allow excellent exposure.[4] Due to being cut into large blocks and slabs, that usually end stored near the quarries, detailed, three-dimensional study of the rock's composition and fossils can be easily done.[5] [6]

Characteristics

The Aurisina limestone has usually a light grey background, sometimes appearing hazelnut in color. It is known for its high purity, compactness, and uniformity. The quarries where it is extracted consist of thick, solid layers of stone. Different varieties of Aurisina marble are distinguished by shades of grey and the distribution, size, and orientation of the organic remains within the stone. Well-known varieties include Aurisina Fiorita, Aurisina Lumachella, Aurisina Chiara, and Roman Stone, among others.[7] [8]

Aurisina marble's chemical composition and physical properties make it ideal for use in architecture, both for structural and decorative purposes, as well as in statues and monuments. The stone is valued for its durability and aesthetic appeal in various applications.[9]

Historical Importance

The Aurisina Limestone has been used since the Roman Republic. Evidence from ancient monuments in Aquileia, a Roman colony founded in 181 BC, suggests that quarrying began in the 1st century BC and continued until the 5th century AD. The stone was widely used in Aquileia for architectural elements, statues, and funerary monuments.[10] [11] The nearby city of Tergeste (modern Trieste) also made extensive use of Aurisina Limestone for public buildings and sculptures. Its use spread across northern Italy, with artifacts found as far as Pavia.

In the post-Roman period, Aurisina marble was notably used in the Mausoleum of Theodoric in Ravenna, built around 520 AD. However, its use declined during the medieval and Baroque periods. In Venice, it was replaced by Istrian stone, but in Trieste, it saw a revival in the 18th century as the city grew following the establishment of the free port by Emperor Charles VI. Notable examples of its use in Trieste include Palazzo Pitteri (1780), the Stock Exchange (1802), and the renovation of the Greek Orthodox Church of San Nicolò dei Greci (1819).[12]

Aurisina Limestone's "golden age" came under the Austro-Hungarian Empire, particularly after the completion of the Southern Railway, connecting Trieste to Vienna. The stone was used in major projects throughout the empire, including the Parliament and Hofburg in Vienna, the State Opera in Budapest, and many buildings in cities like Graz, Munich, and Ljubljana. By 1890, the industry employed over 3,000 workers.

After Trieste's annexation to Italy in 1918, Aurisina Limestone was used in large-scale projects like the Military Shrine of Redipuglia and Milan's Central Station, where 38,000 tons were utilized. During this period, the marble also reached international markets, being used in Egypt and the United States.

In the post-war era, Aurisina Limestone continued to be popular in foreign markets, with notable projects including the Atlanta subway in the U.S., Berlin airport, and La Défense in Paris. In 1989, it was used for Milan Metro's Line 3. More recently, it has featured in prestigious architectural projects, such as Milan's Citylife Shopping District designed by Zaha Hadid and Turin's Lavazza headquarters. Internationally, it is used in projects across Europe, Asia, Australia, and the U.S.[13]

Paleoenvironment

The formation is characterized by lagoonal facies with episodes of higher energy and rare bivalve patch reefs, such as Chondrodonta and Rudists, which are found throughout the entire unit. The lower section, referred to as the "Zolla member" consists of bivalve-rich limestone, including rudists and Chondrodonta, as well as foraminiferal limestone, often interbedded with dolomitized dark wackestones. In the upper part of this member, pelagic limestone with Pythonella fossils is present.[14] [15] The main body of the Aurisina Limestone is characterized by peritidal carbonates, with some dark laminated facies, dominated by rudists, benthic foraminifera, cyanobacteria (Decastronema), and algae (Thaumatoporella).[16] [17] The Paleoenvironment of this unit is accepted to be a succession of emerged and shallow marine carbonate platform settings, including inner lagoons, high-energy shoals, tidal channels, and rudist accumulations[18] The complex dolomitization and silicification in the region may be linked to a Monsoon climate with alternating wet and dry periods. Comparable dolomitization occurs across the Adriatic Platform, particularly in the Middle-Lower Cenomanian.[19]

The Villaggio del Pescatore site revelated bauxite and coal levels in the surrounding sediment indicate a humid climate. This karst region directed most water underground, limiting surface streams. Fossils of Hadrosaurs were found in a 10-meter layer of laminated limestone within a karst depression, unlike other hadrosauroids typically preserved in fluvial or marine deposits.[20] The site also contains remains of small crocodyliforms, fish, and crustaceans, which suggest fluctuating anoxic conditions that preserved the fossils. This depression likely functioned as a cenote, attracting animals seeking water, where some may have drowned and remained preserved in low-oxygen conditions at the sinkhole's base.

Biota

Bivalves

GenusSpeciesLocationMaterialDescriptionImages
Biradiolites[21] [22]
  • B. zucchii[23]
  • B. fissicostatus
  • B. angulosus
  • B. rotundatus
  • B. sp.
  • Cava Romana
  • Nanos Mountain
  • Sistiana[24]
Isolated ShellsRudist of the family Radiolitidae
Bournonia
  • B. excavata
  • B. spp.
  • Cava Romana
  • Nanos Mountain
Isolated ShellsRudist of the family Radiolitidae
Chondrodonta[25] [26]
  • C. joannae
  • C. munsoni
  • Malchina
  • Visogliano
  • Hrusica
  • Nanos Mts
Isolated ShellsScallop of the family Chondrodontidae
Durania[27]
  • D. acuticostata
  • Malchina
  • Visogliano
Isolated ShellsRudist of the family Radiolitidae
Eoradiolites
  • E. fleuriausus
  • E. liratus
  • Malchina
Isolated ShellsRudist of the family Radiolitidae
Gorjanovicia
  • G. lipparinii
  • G. sp.
  • Cava Romana
  • Nanos Mountain
Isolated ShellsRudist of the family Radiolitidae
Hippurites
  • H. nabresinensis
  • Cava Romana
Isolated ShellsRudist of the family Hippuritidae
Katzeria
  • K. hercegovinaensis
  • K. sp.
  • Cava Romana
Isolated ShellsRudist of the family Radiolitidae
Medeella
  • M. zignana
  • Nanos Mountain
Isolated ShellsRudist of the family Radiolitidae
Neithea
  • N. fleuriausiana
  • N. spp.
  • Visogliano
  • Hrusica
  • Nanos Mts
Isolated ShellsScallop of the family Neitheoidae
Paronaites
  • P. zuffardii
  • Malchina
Isolated ShellsRudist of the family Radiolitidae
Praeradiolites
  • P. acutilamellosus
  • P. (cf. Praeradiolites) fleuriausus
  • P. sp.
  • Visogliano
  • Sistiana
Isolated ShellsRudist of the family Radiolitidae
Rajka
  • R. sp.
  • Cava Romana
Isolated ShellsRudist of the family Radiolitidae
Radiolites
  • R. carsicus
  • R. praegalloprovincialis
  • R. peroni
  • R. spp.
  • Visogliano
  • Cava Romana
  • Sistiana
Isolated ShellsRudist of the family Radiolitidae
Radiolitella
  • R. forojuliensis
  • Nanos Mountain
Isolated ShellsRudist of the family Radiolitidae
Requienia
  • R. ssp.
  • Sistiana
Isolated ShellsRudist of the family Requieniidae
Sauvagesia
  • S. sharpei
  • Malchina
  • Visogliano
Isolated ShellsRudist of the family Radiolitidae
Vaccinites
  • V. braciensis
  • V. oppeli
  • V. archiaci
  • V. vredenburgi
  • V. sulcatus
  • Cava Romana
  • Nanos Mountain
Isolated ShellsRudist of the family Hippuritidae

Crustaceans

!Genus!Species!Location!Material!Description!Images
Caridea[29] [30] Indeterminate
  • Villaggio del Pescatore
  • Polazzo
Multiple specimens

Bony Fish

!Genus!Species!Location!Material!Description!Images
AnguilliformesIndeterminatePolazzoSingle complete specimenThis would be the oldest anguilliform found in Italy
Cypriniformes?IndeterminatePolazzoSeveral small complete and incomplete specimensThis determination still needs to be confirmed
EnchodusIndeterminate
  • Polazzo
  • Dobravlje
Several small complete and incomplete specimensA member of Enchodontidae
HolocentridaeIndeterminatePolazzoSeveral small complete and incomplete specimensRepresented mostly by small-sized specimens
ParachanosP. ssp.PolazzoSeveral isolated and articulated remainsA member of Chanidae. The attribution to this genus is dubious
PycnodontiformesIndeterminatePolazzoMultiple complete specimens and isolated teethSome specimens have been referred to the genus Coelodus
RhynchodercetisR. acutissimusPolazzoMultiple specimens, complete and incompleteA member of Dercetidae. The most abundant genus at Polazzo
Sloveniantriacanthus[33] S. saksiŠepulje, east of TomajPMS, VS-001, single specimenA member of Cretatriacanthidae
Teleost indet.Indeterminate
  • Villaggio del Pescatore
  • Polazzo
Multiple specimens, complete and incomplete, as well isolated teeth and scales

Crocodiles

!Genus!Species!Location!Material!Description!Images
Acynodon[34] [35]
  • A.adriaticus
  • A sp.[36]
  • Villaggio del Pescatore
  • Kozina, Kras
MCNST 57248, A partial skeleton in anatomical connection.Multiple referred specimens: MCNST 57032, MCSNT 57031, MCSNT 57245, MCSNT 21.S239, MCSNT 21.S239A small Durophagous crocodrylomorph
CrocodyliaIndeterminate
  • Villaggio del Pescatore
  • Kozina, Kras
MCSNT 57033, possibly a rib; MCSNT 57035, mandibular symphysis; MCSNT 57036, anteroposteriorly long vertebra; MCSNT 57037, fragmented ventral osteodermal surface. Referred Multiple teeth

Pterosaurs

!Genus!Species!Location!Material!Description!Images
PterosauriaIndeterminateVillaggio del PescatoreSpecimen number 13450, an alar metacarpal.

Dinosaurs

!Genus!Species!Location!Material!Description!Image
Coelurosauria[37] IndeterminateKriž, southwest of Tomaj N° BJ 1742 - featherReferral not confirmed
DromaeosauridaeIndeterminateKozina, Kras ACKK-D-8/081, teethResembles other European taxa
HadrosauroideaIndeterminateKozina, Kras ACKK-D-8/20, 121, teethDominant teeth at the locality
Iguanodontidae?IndeterminateKozina, Kras ACKK-D-8/25, 8/04, teethResemble Iguanodontid Teeth, but can belong to a convergently evolved Rhabdodontid
OrnithopodaIndeterminateKozina, Kras ACKK-D-8/21-Z1, teethDifferent from Rhabdodon
Tethyshadros[38] T.insularisVillaggio del PescatoreTwo nearly complete skeletons, a pair of front legs, a left pubis, a cervical vertebra with a still articulated right cervical rib, a complete but highly deformed skull, a series of caudal vertebrae and isolated dorsal rib.A small hadrosauromorph dinosaur
Theropoda indet.[39] IndeterminateVillaggio del PescatoreSC 57030, "arctometatarsalian metatarsal"; Another specimen may represent an epiphysis of a long bone, perhaps of a femur.
TroodontidaeIndeterminateKozina, Kras ACKK-D-8/088, teethTroodontid-alike

Flora

Palynology of the Villaggio del Pescatore section has been studied, dominated by a few gymnosperms and several angiospermous pollen types.

!Genus!Species!Location!Material!Description!Images
AraucaritesA.? sp. KazljeIsolated ConesConifer cones related with Araucariaceae
BrachyphyllumB. spp.PolazzoIsolated Branched ShootsConifer shots related with Araucariaceae or Hirmeriellaceae.
CunninghamitesC. cf. elegansPolazzoIsolated Branched ShootsConifer shots related with Cupressaceae
DicotyledoneaeIndeterminatePolazzoIsolated Leaves
FrenelopsisF. ssp.
  • Villaggio del Pescatore
  • Polazzo
Isolated Branched ShootsA Conifer, member of Hirmeriellaceae. The msot abundant foliar remain locally
Ginkgoales?IndeterminatePolazzoIsolated Leaves
MagnoliaephyllumM. sp.KazljeIsolated LeavesA Flowering Plant, suggested to belong to Magnoliaceae
PagiophyllumP. ssp.
  • Polazzo
  • Kazlje
Isolated Branched ShootsConifer shots related with Araucariaceae or Hirmeriellaceae.
Pinophyta[40] IndeterminateVillaggio del PescatoreIsolated Branched Shoots
Sphenolepiscf.S. spp.PolazzoIsolated ConesConifer cones related with Cupressaceae

See also

References

Notes and References

  1. Consorti . Lorenzo . Arbulla . Deborah . Bonini . Lorenzo . Fabbi . Simone . Fanti . Federico . Franceschi . Marco . Frijia . Gianluca . Pini . Gian Andrea . 2021 . The Mesozoic palaeoenvironmental richness of the Trieste Karst . Geological Field Trips . 13 . 2.2 . 1–40 . 10.3301/gft.2021.06 . 2038-4947. 11368/3009529 . free .
  2. Cucchi . Franco . Forti . Fabio . Finocchiaro . Furio . 1987 . Carbonate surface solution in the classical karst . International Journal of Speleology . 16 . 3/4 . 125–138 . 10.5038/1827-806x.16.3.6 . 0392-6672.
  3. Jurkovšek . Bogdan . Biolchi . Sara . Furlani . Stefano . Kolar-Jurkovšek . Tea . Zini . Luca . Jež . Jernej . Tunis . Giorgio . Bavec . Miloš . Cucchi . Franco . 2016-08-12 . Geology of the Classical Karst Region (SW Slovenia–NE Italy) . Journal of Maps . 12 . sup1 . 352–362 . 10.1080/17445647.2016.1215941 . 2016JMaps..12S.352J . 1744-5647. 11368/2888161 . free .
  4. Sanders . D . 2001 . Burrow-mediated carbonate dissolution in rudist biostromes (Aurisina, Italy): implications for taphonomy in tropical, shallow subtidal carbonate environments . Palaeogeography, Palaeoclimatology, Palaeoecology . 168 . 1–2 . 39–74 . 10.1016/s0031-0182(00)00249-2 . 2001PPP...168...39S . 0031-0182.
  5. Book: Castracane degli Antelminelli, Francesco . Analisi microscopica di un Calcare del Territorio di Spoleto . 1886 . Tipografia delle scienze matematiche e fisiche . Roma. 10.5962/bhl.title.64225 .
  6. Venturini . S. . Tentor . M. . 2010 . La Breccia di Slivia: Una testimonianza di eventi paleotettonici campaniano-maastrichtiani nel Carso Triestino . Natura Nascosta . 41 . 6 . 1–15.
  7. Cancian . Graziano . Princivalle . Francesco . 1989 . Caratteristiche mineralogiche delle grotte del Carso Triestino . International Journal of Speleology . 18 . 3/4 . 111–115 . 10.5038/1827-806x.18.3.3 . 0392-6672.
  8. Book: Martinis, B. . La fragilità del bel paese: geologia dei paesaggi italiani . 2003 . Edizioni Dedalo.
  9. Cucchi . F. . Gerdol . S. . Bertacchi . L. . 1985 . I marmi del Carso triestino . Camera di Commercio, Industria, Artigianato e Agricoltura..
  10. Balcon . S. . Girotto . C. . 2022 . Extraction, processing and trade of Aurisina limestone in the Roman era: the case of the funerary steles of Regio X-Venetia et Histria . The Lenses of the Past Multiscalar Approaches to Archaeology . 49.
  11. Maritan . L. . Mazzoli . C. . Melis . E. . 2003 . A multidisciplinary approach to the characterization of Roman gravestones from Aquileia (Udine, Italy) . Archaeometry . 45 . 3 . 363–374 . 10.1111/1475-4754.00114 . 0003-813X.
  12. Greppi . P. . Bugini . R. . Folli . L. . 2014 . Tecniche e materiali da costruzione nella Milano antica e medievale . LANX. Rivista della Scuola di Specializzazione in Beni Archeologici-Università degli Studi di Milano . 19 . 95–128.
  13. Book: Previato, Caterina . Aurisina Limestone in the Roman Age: From Karst Quarries to the Cities of the Adriatic Basin . 2018 . ASMOSIA XI, Interdisciplinary Studies on Ancient Stone, Proceedings of the XI International Conference of ASMOSIA . http://dx.doi.org/10.31534/xi.asmosia.2015/08.12 . University of Split, Arts Academy in Split; University of Split, Faculty of Civil Engineering, Architecture and Geodesy . 933–939 . 10.31534/xi.asmosia.2015/08.12. 978-953-6617-51-7 .
  14. Consorti . Lorenzo . Arbulla . Deborah . Bonini . Lorenzo . Fabbi . Simone . Fanti . Federico . Franceschi . Marco . Frijia . Gianluca . Pini . Gian Andrea . 2021 . The Mesozoic palaeoenvironmental richness of the Trieste Karst . Geological Field Trips . 13 . 2.2 . 1–40 . 10.3301/gft.2021.06 . 2038-4947. 11368/3009529 . free .
  15. DEBORAH . ARBULLA . 2017 . Ritrovamenti Paleontologici. Il Sito A Dinosauri Del Villaggio Del Pescatore . Atti e Memorie della Commissione Grotte “E. Boegan” . 47 . 87-107.
  16. Previato . Caterina . 2018 . Aurisina Limestone in the Roman Age: from Karst Quarries to the Cities of the Adriatic Basin . ASMOSIA XI, Interdisciplinary Studies on Ancient Stone, Proceedings of the XI International Conference of ASMOSIA . University of Split, Arts Academy in Split; University of Split, Faculty of Civil Engineering, Architecture and Geodesy . 933–939 . 10.31534/xi.asmosia.2015/08.12.
  17. Jurkovšek . Bogdan . Biolchi . Sara . Furlani . Stefano . Kolar-Jurkovšek . Tea . Zini . Luca . Jež . Jernej . Tunis . Giorgio . Bavec . Miloš . Cucchi . Franco . 2016-08-12 . Geology of the Classical Karst Region (SW Slovenia–NE Italy) . Journal of Maps . 12 . sup1 . 352–362 . 10.1080/17445647.2016.1215941 . 1744-5647. 11368/2888161 . free .
  18. Sanders . D . 2001 . Burrow-mediated carbonate dissolution in rudist biostromes (Aurisina, Italy): implications for taphonomy in tropical, shallow subtidal carbonate environments . Palaeogeography, Palaeoclimatology, Palaeoecology . 168 . 1-2 . 39–74 . 10.1016/s0031-0182(00)00249-2 . 0031-0182.
  19. Venturini . S. . Tentor . M. . 2011 . Le rudiste di Malchina conservate presso il Museo della Roca di Monfalcone: un esempio di preservazione di macrofossili calcitici in facies dolomitiche . Natura Nascosta . 42 . 23-37.
  20. Dalla Vecchia . F. M. . 2020 . The unusual tail of Tethyshadros insularis (Dinosauria, Hadrosauroidea) from the Adriatic Island of the European Archipelago . Rivista Italiana di Paleontologia e Stratigrafia . 126 . 3 . 583-628.
  21. Sanders . D . 2001 . Burrow-mediated carbonate dissolution in rudist biostromes (Aurisina, Italy): implications for taphonomy in tropical, shallow subtidal carbonate environments . Palaeogeography, Palaeoclimatology, Palaeoecology . 168 . 1-2 . 39–74 . 10.1016/s0031-0182(00)00249-2 . 0031-0182.
  22. Caffau . Mauro . Pleničar . Mario . 2004-06-30 . Rudist and foraminifer assemblages in the Santonian-Campanian sequence of Nanos Mountain (Western Slovenian) . Geologija . 47 . 1 . 41–54 . 10.5474/geologija.2004.004 . 0016-7789. free .
  23. Caffau . Mauro . Pleničar . Mariop . 1990-06-30 . Biradiolites zucchii n.sp. from the Cava Romana di Aurisina . Geologija . 33 . 1 . 207–213 . 10.5474/geologija.1990.003 . 0016-7789. free .
  24. Caffau . Mauro . Plenicar . Mario . Ogorelec . Bojan . 1998 . Remarks on the morphological variability of Biradiolites angulosus D'Orbigny in a sector of the Trieste Karst (Italy) . Geobios . 31 . 29–36 . 10.1016/s0016-6995(98)80062-4 . 0016-6995.
  25. Venturini . S. . Tentor . M. . 2011 . Le rudiste di Malchina conservate presso il Museo della Roca di Monfalcone: un esempio di preservazione di macrofossili calcitici in facies dolomitiche . Natura Nascosta . 42 . 23-37.
  26. Caffau . Mauro . Pugliese . Nevio . Pleničar . Mario . 1995-12-30 . The development of the mollusc fauna in the Cenomanian of the stratigraphie sequence of Visogliano (Karst of Trieste, Italy) . Geologija . 37/38 . 1 . 87–121 . 10.5474/geologija.1995.003 . 0016-7789. free .
  27. Pons . Jose Maria . Vicens . Enric . Tarlao . Alceo . 2011 . Cenomanian radiolitid bivalves from Malchina, Karst of Trieste, Italy . Cretaceous Research . 10.1016/j.cretres.2011.03.009 . 0195-6671.
  28. Summesberger . H. . Jurkovšek . B. . Kolar-Jurkovšek . T. . 1999 . Upper jaws of Placenticeratidae from the Karst Plateau (Upper Cretaceous, Slovenia). . Annalen des Naturhistorischen Museums in Wien. Serie A für Mineralogie und Petrographie, Geologie und Paläontologie, Anthropologie und Prähistorie . 101 . 3 . 119-122.
  29. Dalla Vecchia . F. M. . Muscio . G. . 1999 . Mesozoic Lagerstaetten of the Friuli-Venezia Giulia region (NE Italy). . Rivista del Museo Civio di Scienzi Naturali "E. Caffi" . 20 . 71–74.
  30. Dalla Vecchia . F.M . Tentor . M. . 2004 . Il Carso 85 milioni di anni fa: Gli straordinari fossili di Polazzo . Gruppo Speleologico Monfalconese A.d.F . 1–75.
  31. Jurkovšek . Bogdan . Kolar-Jurkovšek . Tea . 2007-08-17 . Fossil assemblages from the Upper Cretaceous Komen and Tomaj Limestones of Kras (Slovenia) . Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen . 245 . 1 . 83–92 . 10.1127/0077-7749/2007/0245-0083 . 0077-7749.
  32. Jurkovšek . B. . Kolar-Jurkovšek . T. . 1995 . Upper Cretaceous ray Rhinobatos from the Lipica formation near Dobravlje (Trieste–Komen Plateau, Slovenia) . Annals . 7 . 161-170.
  33. Tyler . J. C. . Križnar . M. . 2013 . A new genus and species, Slovenitriacanthus saksidai, from southwestern Slovenia, of the Upper Cretaceous basal tetraodontiform fish family Cretatriacanthidae (Plectocretacicoidea). . Bollettino del Museo Civico di Storia Naturale di Verona . 37 . 45–56.
  34. DELFINO . MASSIMO . MARTIN . JEREMY E. . BUFFETAUT . ERIC . 2008 . A new species of Acynodon (Crocodylia) from the upper cretaceous (Santonian–Campanian) of Villaggio del Pescatore, Italy . Palaeontology . 51 . 5 . 1091–1106 . 10.1111/j.1475-4983.2008.00800.x . 2008Palgy..51.1091D . 0031-0239.
  35. Marco . Muscioni . Chiarenza . Alfio Alessandro . Delfino . Massimo . Fabbri . Matteo . Milocco . Kevin . Fanti . Federico . 2023 . Acynodon adriaticus from Villaggio del Pescatore (Campanian of Italy): Anatomical and chronostratigraphic integration improves phylogenetic resolution in Hylaeochampsidae (Eusuchia) . Cretaceous Research . 151 . 105631 . 10.1016/j.cretres.2023.105631 . 2023CrRes.15105631M . 0195-6671. 11093/5418 . free .
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  37. Buffetaut . Eric . Jurkovšek . Bogdan . Kolar-Jurkovšek . Tea . 2002 . A fossil feather from the Upper Cretaceous of Kras (Slovenia) . Comptes Rendus Palevol . 1 . 8 . 705–710 . 10.1016/s1631-0683(02)00079-9 . 2002CRPal...1..705B . 1631-0683.
  38. Chiarenza. Alfio Alessandro. Fabbri. Matteo. Consorti. Lorenzo. Muscioni. Marco. Evans. David C.. Cantalapiedra. Juan L.. Fanti. Federico. 2021-12-02. An Italian dinosaur Lagerstätte reveals the tempo and mode of hadrosauriform body size evolution. Scientific Reports. en. 11. 1. 23295. 10.1038/s41598-021-02490-x. 34857789 . 8640049 . 2021NatSR..1123295C . 2045-2322.
  39. Figure 1 Scientific Reports. Scientific Reports . en. 2045-2322.
  40. Web site: Dinosauro Antonio – Museo di Storia Naturale. 2021-12-11. it-IT.