2012 in arthropod paleontology explained

This list of fossil arthropods described in 2012 is a list of new taxa of trilobites, fossil insects, crustaceans, arachnids and other fossil arthropods of every kind that have been described during the year 2012. The list only includes taxa at the level of genus or species.

Arachnids

NameNoveltyStatusAuthorsAgeUnitLocationNotesImages
Ampezzoa[1] Gen. et sp. nov.ValidLindquist & Grimaldi in Schmidt et al.CarnianA mite. The type species is Ampezzoa triassica.
ArchaelagonopsGen. et sp. novValidWunderlichCretaceousBurmese amberA member of Lagonomegopidae. Type species is A. salticoides.
Baltleucauge propinquaSp. novValidWunderlichEoceneBaltic amberEuropeA long-jawed orb weaver
Craspedisia yapchoontecki[2] Sp. nov.ValidPenney et al.MioceneDominican amberA theridiid
Eodoter lonimammillaeSp. novValidWunderlichEoceneBaltic amberEuropeA sac spider
Eohalinobius calefactusSp. novValidWunderlichEoceneBaltic amberEuropeA succinomid
Eohalinobius hiddenseeensisSp. novValidWunderlichEoceneBaltic amberEuropeA succinomid
Eohalinobius patinaSp. novValidWunderlichEoceneBaltic amberEuropeA succinomid
?Eophantes seorsumSp. novValidWunderlichEoceneBaltic amberEuropeA linyphiid
Ephalmator tredecimSp. novValidWunderlichEoceneBaltic amberEuropeAn ephalmatorid
Ero (Succinero) clunis[3] Sp. novValidWunderlichEoceneBaltic amberEurope
Ero (Succinero) gracilitibialisSp. novValidWunderlichEoceneBaltic amberEuropeA pirate spider
Ero (Succinero) vetaSp. novValidWunderlichEoceneBaltic amberEuropeA pirate spider
Geratonephila[4] Gen. et sp. nov.jr synonymPoinar in Poinar & BuckleyEarly CretaceousBurmese amberA nephiline. The type species is G. burmanica.
Moved to Nephila in 2015.
Graea magnocoliSp. novValidWunderlichEoceneBaltic amberEuropeAn orb-weaver spider
HypotheridiosomaGen. et sp. novValidWunderlichCretaceousBurmese amberOriginally classified as a ray spider, but subsequently transferred to the araneoid family Zarqaraneidae. The type species is H. paracymbium.
Korearachne[5] gen et sp novValidSelden et al..Early CretaceousJinju FormationA member of Araneomorphae, possibly a lycosoid. The type species is K. jinju.
LagonoburmopsGen. et sp. novValidWunderlichCretaceousBurmese amberA member of Lagonomegopidae. The type species is L. plumosus.
Leclercera longissipesSp. novJr synonymWunderlichCretaceousBurmese amberA member of Psilodercidae. Originally described as a species of Leclercera, but subsequently transferred to the genus Priscaleclercera.[6]
Leclercera spiculaSp. novJr synonymWunderlichCretaceousBurmese amberA member of Psilodercidae. Originally described as a species of Leclercera, but subsequently transferred to the genus Priscaleclercera.
LeviunguisGen. et sp. novValidWunderlichCretaceousBurmese amberOriginally classified as a ray spider, subsequently transferred to the separate araneoid family Leviunguidae.[7] The type species is L. bruckschi.
Mesobunus dunlopi[8] Sp. nov.ValidGiribet et al.Middle JurassicDaohugou BedsA sclerosomatid harvestman.
MyanlagonopsGen. et sp. novValidWunderlichCretaceousBurmese amberA member of Lagonomegopidae. The type species is M. gracilipes.
OcululoborusGen. et sp. novValidWunderlichCretaceousBurmese amberA member of Uloboridae. The type species is O. curvatus.
Orchestina (Baltorchestina) angulata[9] Nom. novValidWunderlichEoceneBitterfeld amberAn oonopid; a replacement name for Orchestina (Baltorchestina) rectangulata Wunderlich (2011).
Orchestina gappisp novValidSaupe et al..Latest Albian or earliest CenomanianChartense amberAn oonopid
Orchestina rabagensis[10] SpeciesValidSaupe et al.Early AlbianAn oonopid, a species of Orchestina.
Palaeogrosphus jacquesi[11] Sp. novValidLourenço & HenderickxHoloceneMalagasy copalA buthid scorpion.
PalaeoleptonetaGen. et sp. novValidWunderlichCretaceousBurmese amberOriginally described as a member of Leptonetidae, subsequently considered to be a member of Araneomorphae of uncertain affinities by Magalhaes et al..[12] The type species is P. calcar.
Palaeolychas weitschati[13] Sp. nov.ValidLourençoEocene (Kaliningrad Oblast)A buthid scorpion, a species of Palaeolychas.
Palaeotrilineatus[14] Gen. et sp. nov.ValidLourençoEarly CretaceousBurmese amberA scorpion. The type species is P. ellenbergeri.
?Pholcochyrocer baculum[15] Sp. novValidWunderlichCretaceousBurmese amberA member of Pholcochyroceridae, possibly a species of Pholcochyrocer.
Pholcochyrocer pectenSp. novValidWunderlichCretaceousBurmese amberA member of Pholcochyroceridae, a species of Pholcochyrocer.
Piankhi[16] Gen. et sp. nov.ValidDunlop, Bartel & MitovEoceneBaltic amberEuropeA harvestman found in Baltic amber. The type species is P. steineri.
?Poliochera cretacea[17] Sp. novValidWunderlichCretaceousBurmese amberA member of Ricinulei, possibly a species of Poliochera.
Praoppiella[18] Gen. et sp. nov.ValidMiko & Mourek in Miko et al.Early PleistoceneA mite. The type species is P. oanae.
Pseudogarypus synchrotron[19] Sp. novValidHenderickx in Henderickx, Tafforeau & SorianoLate EoceneBaltic AmberA pseudoscorpion
?Psiloderces filiformisSp. novValidWunderlichCretaceousBurmese amberFirst described as possibly species of Psiloderces; transferred to the genus Eopsiloderces.
RhinoppioidesGen. et sp. nov.ValidMiko in Miko et al.Early PleistoceneA mite. The type species is R. quadrituberculatus.
SaetosomaGen. et sp. novValidWunderlichCretaceousBurmese amberA member of Tetrablemmidae. The type species is S. filiembolus.
SamlandicmetaGen. et sp. novValidWunderlichEoceneBaltic amberA long-jawed orb weaver. The type species is S. mutila.
?Scytodes haniSp. novValidWunderlichCretaceousLebanese amberA spider of uncertain affinities. Originally described as a spitting spider, possibly a species of Scytodes; Magalhaes et al. subsequently argued that it cannot be reliably assigned to Scytodoidea.
Succiniropsis runcinataSp. novValidWunderlichEoceneBaltic amberA zoropsid.
?Succinomus gibbosusSp. novValidWunderlichEoceneBaltic amberA succinomid, possibly a species of Succinomus.
Trhypochthonius lopezvallei[20] Sp. nov.ValidArillo, Subías & ShtanchaevaEarly Cretaceous (Albian)A trhypochthoniid mite.
TriasacarusGen. et sp. nov.ValidLindquist & Grimaldi in Schmidt et al.CarnianA mite. The type species is T. fedelei.

Crustaceans

NameNoveltyStatusAuthorsAgeUnitLocationNotesImages
Acanthocythereis inouei[21] Sp. novValidYamaguchi, Mashiba & KamiyaMioceneOsaki FormationAn ostracod, a species of Acanthocythereis.
Acanthophoenicides[22] Gen. et sp. novValidAudo & CharbonnierLate Cretaceous (Cenomanian)Hadjoula LagerstätteA slipper lobster belonging to the subfamily Arctidinae. The type species is A. peterpani.
Acratia candyae[23] Sp. novValidForelLate Permian (Changhsingian)Wujiaping FormationAn acratiid ostracod, a species of Acratia.
AgnocarcinusGen. et sp. novValidBeschin et al.Eocene (Lutetian)A crab belonging to the family Pilumnidae. The type species is A. zannatoi.
AlberticarcinusGen. et sp. novValidBeschin et al.Eocene (Lutetian)A crab belonging to the family Atelecyclidae. The type species is A. eocaenus.
Ammopylocheles robertboreki[24] Sp. novValidFraaije et al.Middle OxfordianA pylochelid, a species of Ammopylocheles.
Amphicytherura grandicribraSp. novValidPuckett & Colin in Puckett, Colin & MitchellLate Cretaceous (Maastrichtian)A schizocytherid ostracod, a species of Amphicytherura.
Amphicytherura occultaSp. novValidPuckett & Colin in Puckett, Colin & MitchellLate Cretaceous (Maastrichtian)An ostracod. Originally described as a species of Amphicytherura; Antonietto et al. (2013) transferred this species to the genus Dinglecythere.[25]
Amphissites gifuensis[26] Sp. novValidTanaka & Yuan in Tanaka et al.Early PermianAn ostracod, a species of Amphissites.
AntonioraninaGen. et comb. novValidVan Bakel et al.Eocene

A raninid crab. The type species is "Cyrtorhina" globosa Beschin, Busulini, De Angeli & Tessier (1988); genus also contains A. fusseli (Blow & Manning, 1996) and A. ripacurtae (Artal & Castillo, 2005). Van Bakel et al. (2012) tentatively assigned "Cyrtorhina" oblonga Beschin, Busulini, De Angeli & Tessier (1988) to the genus Antonioranina as well; however, Karasawa et al. (2014) made it the type species of a separate genus Claudioranina.
Archaeoniscus coreaensis[27] Sp. novValidPark et al.Early CretaceousJinju FormationAn isopod crustacean, a species of Archaeoniscus.
Asciocythere cabbagehillensisSp. novValidPuckett & Colin in Puckett, Colin & MitchellLate Cretaceous (Maastrichtian)A cytherideid ostracod, a species of Asciocythere.
Asymmetricythere semoupathei[28] Sp. novValidSarrYpresian to Lutetian
A trachyleberidid ostracod, a species of Asymmetricythere.
Aurikirkbya miyakeiSp. novValidTanaka & Ono in Tanaka et al.Early PermianAn ostracod, a species of Aurikirkbya.
Aysegulina chapeltonensisSp. novValidPuckett & Colin in Puckett, Colin & MitchellLate Cretaceous (Maastrichtian)A hemicytherid ostracod, a species of Aysegulina.
Aysegulina riominhoensisSp. novValidPuckett & Colin in Puckett, Colin & MitchellLate Cretaceous (Maastrichtian)A hemicytherid ostracod, a species of Aysegulina.
Aysegulina sagittaSp. novValidPuckett & Colin in Puckett, Colin & MitchellLate Cretaceous (Maastrichtian)A hemicytherid ostracod, a species of Aysegulina.
Aysegulina ventrocurvaSp. novValidPuckett & Colin in Puckett, Colin & MitchellLate Cretaceous (Maastrichtian)A hemicytherid ostracod, a species of Aysegulina.
Bairdia adelineaeSp. novValidForelEarly Triassic (Griesbachian)Daye FormationA bairdiid ostracod, a species of Bairdia.
Bairdia fujisanSp. novValidTanaka & Maeda in Tanaka et al.Early PermianAn ostracod, a species of Bairdia.
Bairdia? hubertiSp. novValidForelLate Permian (Changhsingian)Wujiaping FormationA bairdiid ostracod, a possible species of Bairdia.
Bairdia ikeyaiSp. novValidTanaka in Tanaka et al.Early PermianAn ostracod, a species of Bairdia.
Bairdia jeromeiSp. novValidForelLate Permian (Changhsingian) and Early Triassic (Griesbachian)Daye Formation
Wujiaping Formation
A bairdiid ostracod, a species of Bairdia.
Bairdia minoensisSp. novValidTanaka & Ono in Tanaka et al.Early PermianAn ostracod, a species of Bairdia.
Bairdia spindlicaSp. novValidTanaka & Yuan in Tanaka et al.Early PermianAn ostracod, a species of Bairdia.
Balowella crassicostataSp. novValidLuppoldMiddle JurassicAn ostracod, a species of Balowella.
Bathypluma pliocenica[29] Sp. novValidGarassino, Pasini & MariniEarly PlioceneA retroplumid crab, a species of Bathypluma.
BournelyreidusGen. et comb. et sp. novValidVan Bakel et al.Late Cretaceous to Paleocene



A lyreidid crab. The type species is "Hemioon" eysunesensis Collins & Wienberg Rasmussen (1992); according to the authors of its description genus also contains "Raninella" carlilensis Feldmann & Maxey (1980), "Palaeocorystes" laevis Schlüter in von der Marck & Schlüter (1868), "Raninella" manningi Bishop & Williams (2000), "Raninella" oaheensis Bishop (1978) and "Raninella" tridens Roberts (1962), as well as the new species Bournelyreidus teodorii. On the other hand, Karasawa et al. (2014) transferred B. teodorii to the genus Lyreidus, "R." tridens to the genus Macroacaena and "R." manningi to the genus Heus.
Brachycythere jamaicaensisSp. novValidPuckett & Colin in Puckett, Colin & MitchellLate Cretaceous (Maastrichtian)A trachyleberidid ostracod, a species of Brachycythere.
Bretonia[30] gen et comb novValidSchweitzer et al.CenomanianSables du Perche Formationcrab, New genus for "Xanthosia" danielae (Collins and Breton, 2009).
Buntonia bodergatiSp. novValidSarrThanetian to Ypresian
A trachyleberidid ostracod, a species of Buntonia.
Buntonia nanaSp. novValidPuckett & Colin in Puckett, Colin & MitchellLate Cretaceous (Maastrichtian)A trachyleberidid ostracod, a species of Buntonia.
Buntonia vulgarisSp. novValidPuckett & Colin in Puckett, Colin & MitchellLate Cretaceous (Maastrichtian)A trachyleberidid ostracod, a species of Buntonia.
Bythocypris antoniettae[31] Sp. novValidSciutoEarly PleistoceneAn ostracod, a species of Bythocypris.
Bythocythere solisdeus[32] Sp. novValidSciutoEarly PleistoceneA bythocytherid ostracod, a species of Bythocythere.
Calliaxina altavillensis[33] Sp. novValidBeschin & De AngeliEoceneA member of Callianassidae.
Carinokloedenia onusta carinata[34] Subsp. novValidBecker & FrankeDevonian
An ostracod belonging to the group Palaeocopida and the family Beyrichiidae.
Carcinoplax parsensis[35] Sp. novValidVega et al.Miocene (Burdigalian or Langhian)Mishan FormationA goneplacid decapod crustacean, a species of Carcinoplax.
Carpopenaeus garassinoi[36] Sp. novValidCharbonnier in Petit & CharbonnierLate CretaceousSahel Alma Lagerstätte
Cenocorystes bretoniSp. novValidVan Bakel et al.Late Cretaceous (late Cenomanian)A palaeocorystid crab, a species of Cenocorystes.
Cenomanocarcinus cantabricusSp. novValidVan Bakel et al.Early Cretaceous (Albian)A cenomanocarcinid crab, a species of Cenomanocarcinus.
Coleia martinlutheri[37] sp novValidSchweigert & ErnstSinemurianA coleiid polychelid, a species of Coleia.
Cretacolana[38] Gen. et comb. novValidSchweitzer & FeldmannCenomanianA porcelain crab, a new genus for "Porcellana" antiqua A. Milne-Edwards, 1862.
Cretatrizocheles[39] gen et sp novValidFraaije, Klompmaker, & ArtalAlbian or CenomanianA hermit crab. The type species is Cretatrizocheles olazagutiensis.
CromimaiaGen. et comb. novValidBeschin et al.EoceneA crab belonging to the family Majidae. The type species is "Micromaia" meneguzzoi Beschin, Busulini, De Angeli & Tessier (1985); genus also includes "Micromaia" elegans Beschin, Busulini, De Angeli & Tessier (1985).
Cushmanidea tessieriSp. novValidSarrYpresian to LutetianA cushmanideid ostracod, a species of Buntonia.
Cytherelloidea guineacornensis[40] Sp. novValidPuckett in Puckett, Colin & MitchellLate Cretaceous (Maastrichtian)Guinea Corn FormationA cytherellid ostracod, a species of Cytherelloidea.
Cytheropteron eleonoraeSp. novValidSciutoEarly PleistoceneA cytherurid ostracod, a species of Cytheropteron.
Cytheropteron italoiSp. novValidSciutoEarly PleistoceneAn ostracod, a species of Cytheropteron.
Cytheropteron rossanaeSp. novValidSciutoEarly PleistoceneAn ostracod, a species of Cytheropteron.
Daira coccoiSp. novValidBeschin et al.Eocene (Lutetian)A crab belonging to the family Dairidae; a species of Daira.
Dinochelus steeplensis[41] Sp. novValidTshudy & SawardEocene (Ypresian)London ClayA lobster, a species of Dinochelus.
Distefania renefraaijei[42] Sp. novValidKlompmaker, Feldmann & SchweitzerCretaceous (Albian or Cenomanian)A goniodromitid crab, a species of Distefania.
Dolocythere tuberculata[43] Sp. novValidLuppoldMiddle JurassicAn ostracod. Originally described as a species of Dolocythere; Lord, Cabral & Danielopol (2020) transferred this species to the genus Minyocythere.[44]
Ebullitiocaris elatus[45] Sp. novValidWomack et al.CarboniferousA member of Cladocera, a species of Ebullitiocaris.
Eocytheropteron hazeliSp. novValidPuckett & Colin in Puckett, Colin & MitchellLate Cretaceous (Maastrichtian)A cytherurid ostracod, a species of Eocytheropteron.
Eocytheropteron jamaicaensisSp. novValidPuckett & Colin in Puckett, Colin & MitchellLate Cretaceous (Maastrichtian)A cytherurid ostracod, a species of Eocytheropteron.
Eouroptychus[46] Gen. et sp. novValidDe Angeli & CecconEocene (Ypresian)A member of Chirostylidae. The type species is E. montemagrensis.
Eucytherura mitchelliSp. novValidPuckett & Colin in Puckett, Colin & MitchellLate Cretaceous (Maastrichtian)A cytherurid ostracod, a species of Eucytherura.
Eomunidopsis aldoirarensis[47] Sp. novValidKlompmaker et al.Cretaceous (Albian or Cenomanian)A galatheoid, a species of Eomunidopsis.
Eopaguropsis nidiaquilae[48] Sp. novValidFraaije et al.OxfordianA diogenid, a species of Eopaguropsis.
EosymethisGen. et sp. novValidVan Bakel et al.Eocene (Ypresian)Roda FormationA raninid crab. The type species is Eosymethis aragonensis.
Eohexapus orthogoniusSp. novValidBeschin et al.Eocene (Lutetian)A crab belonging to the family Hexapodidae.
Eucorystes iserbytiSp. novValidVan Bakel et al.Early Cretaceous (Albian)A palaeocorystid crab, a species of Eucorystes.
Eucorystes navarrensisSp. novValidVan Bakel et al.Early Cretaceous (Albian)A palaeocorystid crab, a species of Eucorystes.
Faksecarcinusgen et comb novValidSchweitzer et al.Albian or Cenomanian to DanianA crab, a new genus for "Xanthosia" gracilis.
Ferroranina[49] Gen. et comb. et sp. novValidVan Bakel et al.Late Cretaceous (late Cenomanian to late Campanian)[50]



A palaeocorystid crab. The type species is "Notopocorystes" dichrous Stenzel (1945); genus also contains "Notopocorystes" australis Secretan (1964) and "Notopocorystes" fritschi Glaessner (1929), as well as the new species Ferroranina tamilnadu.
FloricythereisGen. et sp. et comb. novPuckett & Colin in Puckett, Colin & MitchellLate Cretaceous (Maastrichtian)
A trachyleberidid ostracod. Genus includes new species Floricythereis exquisita, as well as F. lixula (Crane, 1965) (the type species was not designated).
Galathea mainensis[51] Sp. novValidCeccon & De AngeliMiddle EoceneA species of Galathea.
Gebiacantha tuscia[52] Sp. novValidGarassino, Pasini, De Angeli & Charbonnier in Garassino et al.Early PlioceneAn upogebiid decapod, a species of Gebiacantha.
Geffenina mariebeatriceae[53] Sp. novValidChitnarin in Chitnarin et al.Late Early PermianTak Fa FormationA knoxitid palaeocopidan ostracod, a species of Geffenina.
Geffenina mariebeatriceaeSp. novValidChitnarin in Chitnarin et al.Early PermianPha Nok Khao FormationA knoxitid palaeocopidan ostracod, a species of Geffenina.
Glyphea wiffenae[54] SpeciesValidFeldmann and StilwellLate CretaceousMaungataniwa SandstoneA glypheid lobster, a species of Glyphea.
Glyptopleura mugiensisSp. novValidTanaka & Yuan in Tanaka et al.Early PermianAn ostracod, a species of Glyptopleura.
Goniodromites kubai[55] sp novValidStarzyk, Krzemiska, & KrzemiskiOxfordianA crab, a species of Goniodromites.
Gonioplacoides[56] Gen. et sp. novValidQuayle & CollinsEoceneHeadon Hill FormationA goneplacid. The type species is Gonioplacoides minuta.
Graptocarcinus urbasaensis[57] Sp. novValidVan Bakel et al.Late CretaceousA dynomenid crab, a species of Graptocarcinus.
Grinioneis pachycosta[58] Sp. novValidGuernet et al.Eocene (Lutetian)Paris BasinA trachyleberidid ostracod, a species of Grinioneis.
GrolamaiaGen. et sp. novValidBeschin et al.Eocene (Lutetian)A crab belonging to the subfamily Pisinae. The type species is G. vicariottoi.
Hadrocarcinus[59] gen et comb et sp novValidSchweitzer, Feldmann & LamannaLate CretaceousAntarcticaA necrocarcinid crab, a new genus for "Necrocarcinus" wrighti. The genus also contains "Necrocarcinus" carinatus and a new species Hadrocarcinus tectilacus.
Harbinia alta[60] Sp. novValidAntonietto et al.Aptian or early AlbianSantana FormationAn ostracod crustacean, a species of Harbinia.
HeadonipusGen. et sp. novValidQuayle & CollinsLate EoceneHeadon BedsA hexapodid. The type species is Headonipus tuberculatus.
Hemiparacytheridea exquisitaSp. novValidPuckett & Colin in Puckett, Colin & MitchellLate Cretaceous (Maastrichtian)A paracytherideid ostracod, a species of Hemiparacytheridea.
HispanigalatheaGen. et 2 sp. novValidKlompmaker et al.Cretaceous (Albian or Cenomanian)A galatheoid. Genus contains two species: Hispanigalathea pseudolaevis and H. tuberosa.
Huanghestheria linxiensis[61] Sp. novValidNiu in Zhang et al.Late PermianLinxi FormationA vertexiine lioestheriid spinicaudatan clam shrimp, a species of Huanghestheria.
JoeraninaGen. et comb. et sp. novValidVan Bakel et al.Cretaceous (Albian to early Campanian)








A palaeocorystid crab. The type species is "Corystes" broderipii Mantell (1844). According to the authors of its description genus also contains "Palaeocorystes" harveyi Woodward (1896), "Eucorystes" japonicus Jimbô (1894), "Notopocorystes" paututensis Collins & Wienberg Rasmussen (1992), "Eucorystes" platys Schweitzer & Feldmann (2002) and "Notopocorystes" syriacus Withers (1928), as well as the new species Joeranina gaspari. On the other hand, Karasawa et al. (2014) transferred "P." harveyi, E. platys and "N." paututensis to the genus Eucorystes; the authors also transferred the species "Notopocorystes" xizangensis Wang (1981) to the genus Joeranina.[62]
Juracrista[63] Gen. et 2 sp. novValidRobins, Feldmann & SchweitzerLate Jurassic (Tithonian)Ernstbrunn LimestoneA munidid. The type species is Juracrista perculta; genus also contains Juracrista costaspinosa.
Jurapylocheles iwonaeSp. novValidFraaije et al.Middle OxfordianA pylochelid, a species of Jurapylocheles.
Kinkelinella circumcostataSp. novValidLuppoldMiddle JurassicAn ostracod, a species of Kinkelinella.
Kloedenella poschmanniSp. novValidBecker & FrankeDevonianAn ostracod.
Koryncheiros[64] Gen. et sp. novValidFeldmann et al.Middle or late AnisianGuanling FormationA decapod crustacean; a clytiopsid erymoid. The type species is Koryncheiros luopingensis.
Kroemmelbeincypris[65] Gen. et comb. novValidPoropat & ColinAptianAn ostracod crustacean, a new genus for the species "Hourcqia" symmetrica Krömmelbein & Weber and "Hourcqia" angulata Krömmelbein & Weber.
Langdaia meesookiSp. novValidChitnarin in Chitnarin et al.Late Early to Middle PermianTak Fa FormationA knoxitid palaeocopidan ostracod, a species of Langdaia.
LeiocythereisGen. et sp. novValidPuckett & Colin in Puckett, Colin & MitchellLate Cretaceous (Maastrichtian)A trachyleberidid ostracod. The type species is Leiocythereis polita.
Lepas sattmanni[66] Sp. novValidHarzhauser & SchlöglEarly MioceneEbelsberg FormationA lepadid barnacle, a species of Lepas.
Lessinicarcinus[67] Gen. et comb. novValidDe AngeliEocene[68]
A crab belonging to the family Pilumnidae. The type species is "Titanocarcinus" euglyphos Bittner (1875).
Lessinipagurus[69] Gen. et sp. novValidBeschin et al.Eocene (Lutetian)A hermit crab. Originally assigned to the family Paguridae, subsequently transferred to the separate family Xylopaguridae by Gašparič et al. (2016).[70] The type species is L. granulatus.
Lienenklausicythere[71] Gen. et comb. novValidUffenorde, Monostori & TóthEocene to OligoceneEuropeAn ostracod, a new genus for "Hermanites" acuticosta and "Hermanites" camela; genus also contains a new, yet-unnamed species.
Ljubimovella debilicostatusSp. novValidLuppoldMiddle JurassicAn ostracod, a species of Ljubimovella.
Lobonotus beschini[72] Sp. novValidDe Angeli & ChecchiEocene (Lutetian)A crab belonging to the family Tumidocarcinidae.
Loxoconcha pindarsensisSp. novValidPuckett & Colin in Puckett, Colin & MitchellLate Cretaceous (Maastrichtian)A loxoconchid ostracod, a species of Loxoconcha.
MarylyreidusGen. et comb. novValidVan Bakel et al.Cretaceous (late Albian–early Cenomanian)Pawpaw FormationA lyreidid crab. The type species is "Notopocorystes" punctatus Rathbun (1935).
Megokkos patagoniensis[73] Sp. novValidSchweitzer et al.EoceneRio Turbio FormationA crab, a species of Megokkos.
Mesoparapylochelesgen et sp novValidFraaije, Klompmaker, & ArtalAlbian or CenomanianA hermit crab. The type species is Mesoparapylocheles michaeljacksoni.
Microcheilinella subovataSp. novValidTanaka & Maeda in Tanaka et al.Early PermianAn ostracod, a species of Microcheilinella.
Microcheilinella tsubogawensisSp. novValidTanaka & Ichida in Tanaka et al.Early PermianAn ostracod, a species of Microcheilinella.
Microcoelonella takfaensisSp. novValidChitnarin in Chitnarin et al.Middle PermianTak Fa FormationA coelonellid palaeocopidan ostracod, a species of Microcoelonella.
Microcoelonella takliensisSp. novValidChitnarin in Chitnarin et al.Middle PermianTak Fa FormationA coelonellid palaeocopidan ostracod, a species of Microcoelonella.
Munida deangelii[74] Sp. novValidRobins, Feldmann & SchweitzerLate Oligocene to early MioceneRío Foyel FormationA munidid, a species of Munida.
Myra guriiSp. novValidVega et al.Miocene (Burdigalian or Langhian)Mishan FormationA leucosiid decapod crustacean, a species of Myra.
NavarradromitesGen. et sp. novValidKlompmaker, Feldmann & SchweitzerCretaceous (Albian or Cenomanian)A goniodromitid crab. The type species is Navarradromites pedroartali.
Navarrahomola[75] Gen. et sp. nov.ValidArtal et al.Cretaceous (Albian or Cenomanian)A homolid crab. The type species is Navarrahomola hispanica.
Neocallichirus bacatus[76] Sp. novValidBusulini, Beschin & TessierEocene (Priabonian)A member of Callianassidae.
Neocytherideis labyrinthoideaSp. novValidGuernet et al.Eocene (Lutetian)Paris BasinA neocytherideid ostracod, a species of Neocytherideis.
Neocytheromorpha priscipacificaSp. novValidYamaguchi, Mashiba & KamiyaMioceneOsaki FormationAn ostracod, a species of Neocytheromorpha.
Neurocythere jakovlevae[77] Sp. novValidKolpenskaya in Tesakova et al.Late Jurassic (Early Kimmeridgian)An ostracod, a species of Neurocythere.
Nitotacarcinus antipodesSp. novValidSchweitzer et al.EoceneRio Turbio FormationA crab, a species of Nitotacarcinus.
Nodosculda[78] Gen. et sp. novValidFranţescuEarly Cretaceous (late Albian)A mantis shrimp. The type species is Nodosculda fisherorum.
Noetlingocarcinus messinaiSp. novValidBeschin et al.Eocene (Lutetian)A crab belonging to the superfamily Dromioidea and the family Basinotopidae, a species of Noetlingocarcinus.
Nogarhomola[79] Gen. et sp. novValidDe Angeli & AlbertiEocene (Lutetian)A homolid crab. The type species is Nogarhomola aurorae.
Notopocorystes kerri[80] Sp. novValidLuque et al.Early Cretaceous (late Aptian)Paja FormationA palaeocorystid raninoid crab. Initially described as a species of Notopocorystes, but subsequently reassigned to the genus Joeranina.[81]
NykteripteryxGen. et sp. novValidKlompmaker et al.Cretaceous (Albian or Cenomanian)A galatheoid. The type species is Nykteripteryx rostrata.
Ochtholambrus gambillaraiSp. novValidBusulini, Beschin & TessierEocene (Priabonian)A crab belonging to the family Parthenopidae.
Odontodactylus italicusSp. novValidBeschin et al.Eocene (Lutetian)A mantis shrimp belonging to the family Odontodactylidae; a species of Odontodactylus.
OmmaciriaGen. et comb. novValidBeschin et al.EoceneA crab belonging to the family Majidae. The type species is "Micromaia" mainensis Beschin, Busulini, De Angeli & Tessier (1985).
Orthobairdia jeanlouisiSp. novValidForelEarly Triassic (Griesbachian)Daye FormationA bairdiid ostracod, a species of Orthobairdia.
Ovocytheridea rotundaSp. novValidPuckett & Colin in Puckett, Colin & MitchellLate Cretaceous (Maastrichtian)A cytherideid ostracod, a species of Ovocytheridea.
Paguristes gagnaisoni[82] Sp. novValidGagnaisonMioceneA species of Paguristes.
Pagurus valdagnensisSp. novValidBeschin et al.Eocene (Lutetian)A hermit crab belonging to the family Paguridae, a species of Pagurus.
Palaega picena[83] Sp. novValidPasini & GarassinoMioceneA cirolanid isopod, a species of Palaega.
Palaega pisana[84] Sp. novJunior synonymPasini & GarassinoPlioceneA cirolanid isopod. Originally described as a species of Palaega; considered to be a junior synonym of Bathynomus sismondai (Ristori, 1891) by Hyžný, Pasini & Garassino (2019).[85]
Palaega steatopigia[86] Sp. novNomen dubiumPasini & GarassinoPlioceneA cirolanid isopod. Originally described as a species of Palaega; Hyžný, Pasini & Garassino (2019) considered this species to be a nomen dubium and reassigned it to the genus Bathynomus.
Palaeohomarus pacificus[87] Sp. novValidAguirre-Urreta, Lazo & RawsonLate Hauterivian or early BarremianAgrio FormationA decapod crustacean, a species of Palaeohomarus.
Paracypris keijiSp. novValidGuernet et al.Eocene (Lutetian)Paris BasinA candonid ostracod, a species of Paracypris.
Paranecrocarcinus ballaSp. novValidVan Bakel et al.Late Cretaceous (Cenomanian)A necrocarcinid crab, a species of Paranecrocarcinus.
Periosocypris[88] Gen. et sp. novValidWhatley, Khosla & RathoreLate Cretaceous (Maastrichtian)Lameta FormationA cypridid ostracod. The type species is Periosocypris megistus.
Petrochirus saviiSp. novValidBeschin et al.Eocene (Lutetian)A hermit crab belonging to the family Diogenidae, a species of Petrochirus.
PhrixocythereGen. et sp. novValidPuckett & Colin in Puckett, Colin & MitchellLate Cretaceous (Maastrichtian)A hemicytherid ostracod. The type species is Phrixocythere unionensis.
PlanocarcinusGen. et comb. novValidLuque et al.Early Cretaceous (late Aptian)Paja FormationA necrocarcinid raninoid crab, a new genus for "Dakoticancer" olssoni Rathbun (1937).
Podocallichirus laepaensis[89] Sp. novValidHyžný & MuñizLate MioceneA callianassid ghost shrimp. Originally described as a species of Podocallichirus; subsequently transferred to the genus Balsscallichirus by Hyžný (2016).[90]
Protobuntonia waroreiSp. novValidSarrPaleocene to Ypresian
A trachyleberidid ostracod, a species of Buntonia.
Pterygocythereis babinotiSp. novValidPuckett & Colin in Puckett, Colin & MitchellLate Cretaceous (Maastrichtian)A trachyleberidid ostracod, a species of Pterygocythereis.
Pylochelitergites[91] Gen. et 2 sp. novValidFraaije et al.Oxfordian to Kimmeridgian
A pylochelid hermit crab. Genus contains two species: P. westerbergensis (the type species) and P. gelasinus.
Quasilaeviranina minusculaSp. novValidBeschin et al.Eocene (Lutetian)A crab belonging to the family Raninidae, a species of Quasilaeviranina.
Raninoides rioturbiensisSp. novValidSchweitzer et al.EoceneRio Turbio FormationA crab, a species of Raninoides.
Rhinolambrus elongatusSp. novValidBeschin et al.Eocene (Lutetian)A crab belonging to the family Parthenopidae; a species of Rhinolambrus.
RistoriaGen. et comb. novValidGarassino, Pasini, De Angeli & Charbonnier in Garassino et al.Early PlioceneA leucosiid crab, a new genus for "Ilia" pliocaenica Ristori, 1891.
Samarella soneiSp. novValidChitnarin in Chitnarin et al.Middle PermianTak Fa FormationA paraparchitid palaeocopidan ostracod, a species of Samarella.
Samarella viscusformaSp. novValidChitnarin in Chitnarin et al.Early to Middle PermianPha Nok Khao Formation
Tak Fa Formation
A paraparchitid palaeocopidan ostracod, a species of Samarella.
Sargentina chantarameeiSp. novValidChitnarin in Chitnarin et al.Early to Middle PermianPha Nok Khao Formation
Tak Fa Formation
A knoxitid palaeocopidan ostracod, a species of Sargentina.
Scalpellum paratethyianumSp. novValidHarzhauser & SchlöglEarly MioceneLakšárska Nová Ves FormationA scalpellid barnacle, a species of Scalpellum.
Schizoptocythere nanaSp. novValidPuckett & Colin in Puckett, Colin & MitchellLate Cretaceous (Maastrichtian)A trachyleberidid ostracod, a species of Schizoptocythere.
Schizoptocythere pinnaSp. novValidPuckett & Colin in Puckett, Colin & MitchellLate Cretaceous (Maastrichtian)A trachyleberidid ostracod, a species of Schizoptocythere.
Schuleridea exiguaSp. novValidPuckett & Colin in Puckett, Colin & MitchellLate Cretaceous (Maastrichtian)A cytherideid ostracod, a species of Schuleridea.
Semicytherura cariouiSp. novValidSarrYpresian to LutetianA cytherurid ostracod, a species of Semicytherura.
Sineruga[92] gen et sp novValid
  • Perrier
Silurian (uppermost Ludlow to lower Pridoli)Le Val Formationmyodocope ostracod. type species is Sineruga insolita.
Southcavea (Saxosouthcavea)Subgen. et 3 sp. novValidLuppoldMiddle JurassicAn ostracod, a subgenus of Southcavea containing three new species: Southcavea (Saxosouthcavea) extendata, Southcavea (Saxosouthcavea) bartenbrandi and Southcavea (Saxosouthcavea) malzi (with two subspecies: S. (S.) malzi malzi and S. (S.) malzi reticulata).
SpinicytherideaGen. et sp. novValidPuckett & Colin in Puckett, Colin & MitchellLate Cretaceous (Maastrichtian)
A cytherideid ostracod. The type species is Spinicytheridea compta from Jamaica; genus also includes a second, unnamed species from the Maastrichtian Ocozocoautla Formation of Chiapas, Mexico.
SpinirostrimaiaGen. et comb. novValidBeschin et al.Eocene
A crab belonging to the family Majidae. The type species is "Micromaia" margaritata Fabiani (1910).
Stagmacaris adielklompmakeriSp. novValidFraaije et al.Early late KimmeridgianA pylochelid hermit crab, a species of Stagmacaris.
Stagmacaris biarcusSp. novValidFraaije et al.Middle OxfordianA pylochelid hermit crab, a species of Stagmacaris.
Steorrosiagen et comb novValidSchweitzer et al.Late Albian to CenomanianCrab, New genus for "Xanthosia" aspera. Other species S. pawpawensis and S. reidi.
StyracocarcinusGen. et comb. novValidSchweitzer & FeldmannLate Cretaceous (Senonian)A tumidocarcinid decapod, a new genus for "Titanocarcinus" meridionalis Secretan, 1961.
SuperpendicythereGen. et sp. novValidLuppoldMiddle JurassicAn ostracod. The type species is S. binodosa.
SymethoidesGen. et sp. novValidVan Bakel et al.Paleocene (early Danian)Tinton FormationA raninid crab. The type species is Symethoides monmouthorum.
Szaboa deppermanni[93] Sp. novValidFraaije et al.OligoceneRatzeburg FormationA matutid crab, a species of Szaboa.
Thalamita italicaSp. novValidGarassino, Pasini, De Angeli & Charbonnier in Garassino et al.Early PlioceneA portunid crab, a species of Thalamita.
TridactylastacusGen. et sp. novValidFeldmann et al.Middle or late AnisianGuanling FormationA decapod crustacean; a glypheoid belonging to the family Litogastridae.[94] The type species is Tridactylastacus sinensis.
TurbiocheirGen. et sp. novDisputedSchweitzer et al.EoceneRio Turbio Formation

[95]
A thalassinidean. The type species is Turbiocheir minutospinata. Hyžný et al. (2020) considered the genus Turbiocheir to be a junior synonym of the genus Anacalliax, creating a new combination Anacalliax minutospinata.[96]
VecticallichirusGen. et sp. novValidQuayle & CollinsLate Eocene and OligoceneHeadon BedsA callichirid ghost shrimp. The type species is Vecticallichirus abditus. Hyžný (2020) considered V. abditus to be a junior synonym of "Callianassa" batei Woodward (1869), resulting in a new combination Vecticallichirus batei.[97]
VegaraninaGen. et comb. novValidVan Bakel et al.Late Cretaceous (Campanian–Maastrichtian)


A raninid crab. The type species is "Lophoranina" precocious Feldmann, Vega, Tucker, García-Barrera & Avendaño (1996).
Vernoniella tenuiporaSp. novValidLuppoldMiddle JurassicAn ostracod, a species of Vernoniella.
ViaiaGen. et sp. nov.ValidArtal et al.Cretaceous (Albian or Cenomanian)A viaiid glaessneropsoid crab. The type species is Viaia robusta.
YunnanopalinuraGen. et sp. novValidFeldmann et al.Middle or late AnisianGuanling FormationA decapod crustacean; a spiny lobster. The type species is Yunnanopalinura schrami.

Eurypterids

NameNoveltyStatusAuthorsAgeUnitLocationNotesImages
Adelophthalmus dubius[98] Sp. novValidShpinevMiddle DevonianA eurypterid, a species of Adelophthalmus.
Adelophthalmus kamyshtensisSp. novValidShpinevMiddle DevonianA eurypterid, a species of Adelophthalmus.
Drepanopterus odontospathus[99] Sp. nov ValidLamsdellEarly DevonianA eurypterid.
Nanahughmilleria notosibirica[100] Sp. novValidShpinevDevonianA eurypterid, a species of Nanahughmilleria.
Parahughmilleria longaSp. novValidShpinevDevonianA eurypterid, a species of Parahughmilleria.
Stylonuroides orientalisSp. novValidShpinevDevonianA eurypterid, a species of Stylonuroides.

Trilobites

NameNoveltyStatusAuthorsAgeUnitLocationNotesImages
Abanomocarella[101] Gen. et sp. et comb. novValidYuan et al.CambrianChanghia FormationA member of Ptychopariida belonging to the family Anomocarellidae. Genus includes new species A. quadrata, as well as A. humilis (Resser & Endo in Endo & Resser, 1937) and A. pyriformis (Resser & Endo in Endo & Resser, 1937).
Acmarhachis kindlei[102] Sp novValidWestrop & EoffLate CambrianCow Head GroupAn agnostid, a species of Acmarhachis.
Acmarhachis whittingtoniSp novValidWestrop & EoffLate CambrianCow Head GroupAn agnostid, a species of Acmarhachis.
Angsiduoa jiulongshanensisSp. novValidYuan et al.CambrianChanghia FormationA member of Ptychopariida belonging to the family Ignotogregatidae.
Angsiduoa shanxiensisSp. novValidYuan et al.CambrianChanghia FormationA member of Ptychopariida belonging to the family Ignotogregatidae.
Angsiduoa transversaSp. novValidYuan et al.CambrianChanghia FormationA member of Ptychopariida belonging to the family Ignotogregatidae.
Anomocarella? antiquaSp. novValidYuan et al.CambrianChanghia FormationA member of Ptychopariida belonging to the family Anomocarellidae.
Anomocarella jiulongshanensisSp. novValidYuan et al.CambrianChanghia FormationA member of Ptychopariida belonging to the family Anomocarellidae.
Anomocarella similisSp. novValidYuan et al.CambrianChanghia FormationA member of Ptychopariida belonging to the family Anomocarellidae.
Aponileus aasei[103] Sp novValidAdrain & McAdamsEarly Ordovician (late Floian)Fillmore FormationA bathyurid trilobite, a species of Aponileus.
Aponileus belkaaeSp novValidAdrain & McAdamsEarly Ordovician (late Floian)Wah Wah FormationA bathyurid trilobite, a species of Aponileus.
Aponileus laikaaeSp novValidAdrain & McAdamsEarly Ordovician (late Floian)Fillmore FormationA bathyurid trilobite, a species of Aponileus.
Aponileus strelkaaeSp novValidAdrain & McAdamsEarly Ordovician (late Floian)Wah Wah FormationA bathyurid trilobite, a species of Aponileus.
Aponileus ugolekaeSp novValidAdrain & McAdamsEarly Ordovician (late Floian)Wah Wah FormationA bathyurid trilobite, a species of Aponileus.
Aponileus? veterokaeSp novValidAdrain & McAdamsEarly Ordovician (late Floian)Wah Wah FormationA bathyurid trilobite, a possible species of Aponileus.
Archaeagnostus typicusSp. novValidYuan et al.CambrianChanghia FormationA member of the family Peronopsidae.
Archegonus (Phillibole) nehdenensis africanus[104] Subsp. novValidHahn, Müller & BeckerCarboniferous (Mississippian)A member of Phillipsiidae belonging to the subfamily Archegoninae, a subspecies of Archegonus nehdenensis.
Asaphellus cuervoae[105] Sp. novDisputedCorbacho & López-SorianoOrdovician (Floian)Upper Fezouata FormationA member of Asaphidae, a species of Asaphellus. Gutiérrez-Marco et al. (2017) considered this species to be likely based on poorly preserved and probably retouched specimens of Asaphellus stubbsi.[106]
Astycoryphe arduinnae[107] Sp novValidVan Viersen, van Rossum & PrescherMiddle DevonianA member of Proetida.
Australoscutellum iota[108] sp novValidHolloway and LaneSilurianA scutelluid trilobite, a species of Australoscutellum.
Australoscutellum micropssp novValidHolloway and LaneSilurianA scutelluid trilobite, a species of Australoscutellum.
Australoscutellum pulexsp novValidHolloway and LaneSilurianA scutelluid trilobite, a species of Australoscutellum.
Australoscutellum tricasp novValidHolloway and LaneSilurianA scutelluid trilobite, a species of Australoscutellum.
Baltagnostus yaojiayuensisSp. novValidYuan et al.CambrianChanghia FormationA member of the family Diplagnostidae.
Baojingia (Eobaojingia)Subgen. et 2 sp. novValidYuan et al.CambrianChanghia FormationA member of the family Lisaniidae. The subgenus includes Baojingia (Eobaojingia) dubia and Baojingia (Eobaojingia) ezhuangensis, as well as Baojingia miaogouensis Mong in Zhou et al. (1977).
Borenoriagen et sp novValidHolloway and LaneSilurianA scutelluid trilobite. The type species is Borenoria cirrita.
Bumastoides graffhami[109] Sp novValidCarlucci et al.OrdovicianAn illaenid trilobite, a species of Bumastoides.
Bumastoides kimmswickensisSp novValidCarlucci et al.OrdovicianAn illaenid trilobite, a species of Bumastoides.
Bumastoides moundensisSp novValidCarlucci et al.OrdovicianAn illaenid trilobite, a species of Bumastoides.
Calymene harringtoni[110] Sp. novValidTortello in Tortello et al.Early SilurianVargas Peña FormationA species of Calymene.
Chlupacula (Chlupacula) erfoudensisSp. novValidHahn, Müller & BeckerCarboniferous (Mississippian)A member of Phillipsiidae belonging to the subfamily Archegoninae, a species of Chlupacula.
Crepicephalina exsculptaSp. novValidYuan et al.CambrianChanghia Formation
Crepicephalina obscuraSp. novValidYuan et al.CambrianChanghia Formation
Crepicephalina pulchraSp. novValidYuan et al.CambrianChanghia Formation
Crepicephalina? raraSp. novValidYuan et al.CambrianChanghia Formation
Crepicephalina woqishanensisSp. novValidYuan et al.CambrianChanghia Formation
Cummingella boikoi[111] sp novValidMychkoSakmarianA phillipsiid, a species of Cummingella.
Cyrtoproetus (Crassibole) acropsSp. novValidHahn, Müller & BeckerCarboniferous (Mississippian)A member of Phillipsiidae belonging to the subfamily Archegoninae, a species of Cyrtoproetus.
Dalejeproetus owensi[112] Sp novValidJohnson & ForteyEarly Devonian (Pragian)Ihandar FormationA proetid trilobite, a species of Dalejeproetus.
Dalejeproetus sagaouiiSp novValidJohnson & ForteyEarly Devonian (Pragian)Ihandar FormationA proetid trilobite, a species of Dalejeproetus.
DameselloidesGen. et sp. novValidYuan et al.CambrianChanghia FormationA member of the family Damesellidae. Genus includes new species D. sinensis.
Danjiangella shandongensisSp. novValidYuan et al.CambrianChanghia Formation
Dechenelloides tazoultensisSp. novValidHahn, Müller & BeckerCarboniferous (Mississippian)A member of Phillipsiidae belonging to the subfamily Archegoninae, a species of Dechenelloides.
Decoroscutellum planessp novValidHolloway and LaneSilurianA scutelluid trilobite, a species of Decoroscutellum.
Derikaspis conicusSp. novValidYuan et al.CambrianChanghia Formation
Derikaspis ruichengensisSp. novValidYuan et al.CambrianChanghia Formation
Diacoryphe (Archaeocoryphe) maiderensisSp. novValidHahn, Müller & BeckerCarboniferous (Mississippian)A member of Phillipsiidae belonging to the subfamily Archegoninae, a species of Diacoryphe.
Dindymene whittingtoni[113] Sp novValidOwen & BrutonLate OrdovicianA species of Dindymene.
Dorypyge cylindricaSp. novValidYuan et al.CambrianChanghia Formation
Dorypyge zhangxianensisSp. novValidYuan et al.CambrianChanghia Formation
Dunopyge seiberti[114] Sp. novValidBasse & MüllerEarly DevonianSeifen BedsA member of Acastidae belonging to the subfamily Asteropyginae, a species of Dunopyge.
Eilura trapezialisSp. novValidYuan et al.CambrianChanghia Formation
Elaphraella? taebaeksanensis[115] Sp. novValidPark & ChoiCambrian (middle Furongian)Sesong FormationA member of the family Shumardiidae.
Emmrichella punctataSp. novValidYuan et al.CambrianChanghia FormationA member of the family Ptychopariidae.
EokoldiniaGen. et sp. novValidYuan et al.CambrianChanghia FormationA member of the family Plethopeltidae. Genus includes new species E. porifera.
EomapaniaGen. et sp. novValidYuan et al.CambrianChanghia FormationA member of the family Mapaniidae. Genus includes new species E. subquadrata.
Eoscutellum annosumsp novValidHolloway and LaneSilurianA scutelluid trilobite, a species of Eoscutellum.
Eymekops pristinaSp. novValidYuan et al.CambrianChanghia Formation
Eymekops transversaSp. novValidYuan et al.CambrianChanghia Formation
Grandioculus macroculusSp. novValidYuan et al.CambrianChanghia Formation
Grandioculus (Protohedina) quadraticephalaSp. novValidYuan et al.CambrianChanghia Formation
Gumunsoia[116] Gen. et sp. novValidPark, Sohn & ChoiCambrian (middle Furongian)Sesong FormationA member of Leiostegioidea belonging to the family Kaolishaniidae. Genus includes new species G. triangularis.
GushanellaGen. et 2 sp. novValidYuan et al.CambrianChanghia FormationA member of the family Solenopleuridae. Genus includes new species G. quadrata and G. truncata.
Heyelingella crassaSp. novValidYuan et al.CambrianChanghia Formation
Honania poriferaSp. novValidYuan et al.CambrianChanghia Formation
Huayuanaspis angustilimbataSp. novValidYuan et al.CambrianChanghia Formation
Idioura brevispinaSp. novValidYuan et al.CambrianChanghia Formation
Illaenoscutellum tryosp novValidHolloway and LaneSilurianA scutelluid trilobite, a species of Illaenoscutellum.
Inouyella raraSp. novValidYuan et al.CambrianChanghia Formation
Inouyella shiliuyuanensisSp. novValidYuan et al.CambrianChanghia Formation
Inouyia (Bulbinouyia)Subgen. novValidYuan et al.CambrianChanghia FormationA member of the family Inouyiidae. The subgenus includes Inouyia kangzhuangensis (Zhang & Wang, 1985) and I. lubrica (Zhou in Zhou et al., 1982).
Inouyia latilimbataSp. novValidYuan et al.CambrianChanghia Formation
Iranoleesia pustulosaSp. novValidYuan et al.CambrianChanghia Formation
Iranoleesia (Proasaphiscina) rogosaSp. novValidYuan et al.CambrianChanghia Formation
Japonoscutellum borenorensesp novValidHolloway and LaneSilurianA scutelluid trilobite, a species of Japonoscutellum.
Japonoscutellum crassumsp novValidHolloway and LaneSilurianA scutelluid trilobite, a species of Japonoscutellum.
Japonoscutellum diascetumsp novValidHolloway and LaneSilurianA scutelluid trilobite, a species of Japonoscutellum.
Japonoscutellum gephyricumsp novValidHolloway and LaneSilurianA scutelluid trilobite, a species of Japonoscutellum.
Japonoscutellum guttulatumsp novValidHolloway and LaneSilurianA scutelluid trilobite, a species of Japonoscutellum.
Japonoscutellum serangicumsp novValidHolloway and LaneSilurianA scutelluid trilobite, a species of Japonoscutellum.
Japonoscutellum strabosp novValidHolloway and LaneSilurianA scutelluid trilobite, a species of Japonoscutellum.
Jiulongshania shandongensisSp. novValidYuan et al.CambrianChanghia FormationA member of the family Diceratocephalidae.
Koptura (Eokoptura)Subgen. et sp. novValidYuan et al.CambrianChanghia FormationA member of the family Anomocaridae. The subgenus includes new species Koptura (Eokoptura) bella, as well as Koptura quadrata (Resser & Endo in Endo & Resser, 1937).
Koptura intermediaSp. novValidYuan et al.CambrianChanghia FormationA member of the family Anomocaridae.
Kosovopeltis trepidasp novValidHolloway and LaneSilurianA scutelluid trilobite, a species of Kosovopeltis.
Kosovopeltis xynonsp novValidHolloway and LaneSilurianA scutelluid trilobite, a species of Kosovopeltis.
Lepidoproetus lahceniSp novValidJohnson & ForteyEarly Devonian (Pragian)Ihandar FormationA proetid trilobite, a species of Lepidoproetus.
Lepidoproetus maharchianusSp novValidJohnson & ForteyEarly Devonian (Pragian)Ihandar FormationA proetid trilobite, a species of Lepidoproetus.
Lepidoproetus splendensSp novValidJohnson & ForteyEarly Devonian (Pragian)Ihandar FormationA proetid trilobite, a species of Lepidoproetus.
Liowutaishania laevigataSp. novValidYuan et al.CambrianChanghia Formation
Lisania ellipticaSp. novValidYuan et al.CambrianChanghia Formation
Longiproetus meiopsifiusSp novValidVan Viersen, van Rossum & PrescherMiddle DevonianA member of Proetida.
LunaniaGen. novJunior homonymYuan et al.CambrianChanghia FormationA member of the family Menocephalidae. The generic name is preoccupied by Lunania Chow (1957).
Maotunia transversaSp. novValidYuan et al.CambrianChanghia Formation
Mapania (Mapanella)Subgen. et 2 sp. novValidYuan et al.CambrianChanghia FormationA member of the family Mapaniidae. The subgenus includes new species M. (Mapanella) transversa and M. (Mapanella) prisca, as well as Mapania blackwelderi (Resser & Endo in Endo & Resser, 1937).
Mapania subcylindricaSp. novValidYuan et al.CambrianChanghia Formation
Mapanopsis conicusSp. novValidYuan et al.CambrianChanghia Formation
Maurotarion megacephalum[117] Sp. nov.ValidRustán & VaccariEarly Devonian (Pragian)Talacasto FormationAn otarionine aulacopleurid, a species of Maurotarion.
Megagraulos similisSp. novValidYuan et al.CambrianChanghia Formation
MenocephalinaGen. et sp. novValidYuan et al.CambrianChanghia FormationA member of the family Menocephalidae. Genus includes new species M. longispinosa.
Menocephalites (Hutoushania)Subgen. et 2 sp. novValidYuan et al.CambrianChanghia FormationA member of the family Menocephalidae. The subgenus includes Menocephalites (Hutoushania) triangulata and M. (Hutoushania) zaozhuangansis.
Menocephalites porosusSp. novValidYuan et al.CambrianChanghia Formation
Menocephalites primitivusSp. novValidYuan et al.CambrianChanghia Formation
Menocephalites simplexSp. novValidYuan et al.CambrianChanghia Formation
Metanomocarella primitivaSp. novValidYuan et al.CambrianChanghia Formation
Micagnostus semicircularisSp. novValidYuan et al.CambrianChanghia Formation
Mirrabookagen et sp novValidHolloway and LaneSilurianA scutelluid trilobite. The type species is Mirrabooka harryi.
Monkaspis conicaSp. novValidYuan et al.CambrianChanghia Formation
Ohleum magreani[118] Sp novValidTaghon, Bonino & MottequinDevonian (Eifelian)Jemelle FormationA member of Lichida belonging to the family Lichidae.
Pagetides qianensis[119] Sp novValidZhang & ClarksonCambrianAn eodiscoid trilobite, a species of Pagetides.
Paleonelsonia granulosaSp. novValidYuan et al.CambrianChanghia FormationA member of the family Ptychopariidae.
Paleonelsonia sulcataSp. novValidYuan et al.CambrianChanghia FormationA member of the family Ptychopariidae.
Paracoosia langgongsiensisSp. novValidYuan et al.CambrianChanghia FormationA member of the family Anomocaridae.
Paralevisia subglobosaSp. novValidYuan et al.CambrianChanghia Formation
Parayujinia pulchraSp. novValidYuan et al.CambrianChanghia Formation
Peishania abnormisSp. novValidYuan et al.CambrianChanghia Formation
Peishania obscuraSp. novValidYuan et al.CambrianChanghia Formation
Peronopsis jiulongshanensisSp. novValidYuan et al.CambrianChanghia Formation
Peronopsis universalisSp. novValidYuan et al.CambrianChanghia Formation
Persia[120] Gen. et sp. novJunior homonymLerosey-AubrilCarboniferous (Tournaisian)The type species is Persia praecox. The generic name is preoccupied by Persia Repin (1996); Lerosey-Aubril & Deshmukh (2022) coined a replacement name Persiax.[121]
Pilletina seifenensisSp. novValidBasse & MüllerEarly DevonianSeifen BedsA member of Acastidae belonging to the subfamily Asteropyginae, a species of Pilletina.
Pingluia granulosaSp. novValidYuan et al.CambrianChanghia Formation
Plectocrania elegansSp. novValidYuan et al.CambrianChanghia Formation
Plectocrania intermediaSp. novValidYuan et al.CambrianChanghia Formation
Plesigangderria latilimbataSp. novValidYuan et al.CambrianChanghia Formation
Podoliproetus mirdaniSp novValidJohnson & ForteyEarly Devonian (Pragian)Ihandar FormationA proetid trilobite, a species of Podoliproetus.
Podoliproetus sinespinaSp novValidJohnson & ForteyEarly Devonian (Pragian)Ihandar FormationA proetid trilobite, a species of Podoliproetus.
PraeymekopsGen. et sp. et comb. novValidYuan et al.CambrianChanghia FormationA member of the family Proasaphiscidae. Genus includes new species P. minutus, as well as P. sinensis (Mansuy, 1916).
Proasaphiscus laiwuensisSp. novValidYuan et al.CambrianChanghia Formation
Proetina ihamadiiSp novValidJohnson & ForteyEarly Devonian (Pragian)Ihandar FormationA proetid trilobite, a species of Proetina.
Pseudoolenoides fossilmountainensis[122] Sp novValidAdrain, McAdams & KarimMiddle OrdovicianKanosh FormationA bathyurid, a species of Pseudoolenoides.
Pseudoolenoides oilcreekensisSp novValidAdrain, McAdams & KarimMiddle OrdovicianOil Creek FormationA bathyurid, a species of Pseudoolenoides.
Pseudoolenoides pogonipensisSp novValidAdrain, McAdams & KarimMiddle OrdovicianKanosh FormationA bathyurid, a species of Pseudoolenoides.
PseudopeishaniaGen. et 3 sp. et comb. novValidYuan et al.CambrianChanghia FormationA member of Ptychopariida belonging to the family Anomocarellidae. Genus includes new species P. brevis, P. similis and P. woqishanensis, as well as P. angustata (Endo, 1944).
Pseudoperonopsis elegansSp. novValidYuan et al.CambrianChanghia Formation
Pseudophillipsia parviziiSp. novValidLerosey-AubrilPermianA species of Pseudophillipsia sensu lato.
Pseudotaitzuia angustilimbataSp. novValidYuan et al.CambrianChanghia FormationA member of Asaphida belonging to the family Ordosiidae.
Pseudotaitzuia raraSp. novValidYuan et al.CambrianChanghia FormationA member of Asaphida belonging to the family Ordosiidae.
Pudoproetus zhoraeSp. novValidHahn, Müller & BeckerCarboniferous (Mississippian)A member of Proetidae, a species of Pudoproetus.
Raragnostus intermediusSp. novValidYuan et al.CambrianChanghia Formation
Raymondaspis grandigena[123] sp novValidStein & BergströmMiddle OrdovicianA styginid, a species of Raymondaspis.
Reediella mitchelli tafilaltensisSubsp. novValidHahn, Müller & BeckerCarboniferous (Mississippian)A member of Phillipsiidae belonging to the subfamily Bollandiinae, a subspecies of Reediella mitchelli.
RuichengocephalusGen. et sp. novValidYuan et al.CambrianChanghia FormationA member of the family Ptychopariidae. Genus includes new species R. typicus.
Saimachia yichangensisSp. novValidYuan et al.CambrianChanghia Formation
Schopfaspis? graciai[124] sp novValidEsteve et al..Middle CambrianAn alokistocarid, possibly a species of Schopfaspis.
Sdzuyella conicaSp. novValidYuan et al.CambrianChanghia FormationA member of Ptychopariida belongig to the family Holocephalidae.
Securifronsgen et comb novValidHolloway and LaneOrdovician to SilurianScutelluid trilobite. species- S. borealis, S. cunctatum, S. major and ?S. umbonatus.
ShiliuyuaniaGen. et 2 sp. novValidYuan et al.CambrianChanghia FormationA member of the family Crepicephalidae. Genus includes new species S. cylindrica and S. quadrata.
Solenoparia zhangxianensisSp. novValidYuan et al.CambrianChanghia Formation
Solenoparina bellaSp. novValidYuan et al.CambrianChanghia Formation
Sphaerophthalmus arcus[125] Sp novValidHøyberget & BrutonCambrian (Furongian)Alum Shale FormationA member of the family Olenidae.
Spiniscabrella albertii[126] Sp. novValidMüller & FrankeDevonianA member of the family Homalonotidae.
Sudanomocarina ellipticaSp. novValidYuan et al.CambrianChanghia Formation
Sudanomocarina hutoushanensisSp. novValidYuan et al.CambrianChanghia Formation
Symphysops stevaninae[127] Sp. novValidLópez-Soriano & CorbachoLate OrdovicianLower Ktaoua FormationA member of Cyclopygidae, a species of Symphysops.
Szeaspis (Pseudoszeaspis)Subgen. et sp. novValidYuan et al.CambrianChanghia FormationA member of Ptychopariida belonging to the family Anomocarellidae. The subgenus includes new species Szeaspis (Pseudoszeaspis) conicus, as well as Szeaspis huananensis Peng, Babcock & Lin (2004) and Szeaspis pustulosa (Resser & Endo in Endo & Resser, 1937).
Szeaspis crassusSp. novValidYuan et al.CambrianChanghia FormationA member of Ptychopariida belonging to the family Anomocarellidae.
Taitzehoia huolianzhaiensisSp. novValidYuan et al.CambrianChanghia FormationA member of the family Damesellidae.
Teinistion jiulongshanenseSp. novValidYuan et al.CambrianChanghia FormationA member of the family Damesellidae.
Teinistion quadratumSp. novValidYuan et al.CambrianChanghia FormationA member of the family Damesellidae.
Teinistion yangjiazhuangenseSp. novValidYuan et al.CambrianChanghia FormationA member of the family Damesellidae.
Tomagnostus (Paratomagnostus)Subgen. et sp. novValidYuan et al.CambrianChanghia FormationA member of the family Ptychagnostidae. The subgenus includes Tomagnostus (Paratomagnostus) ezhuangensis.
Trachoparia elongataSp. novValidYuan et al.CambrianChanghia Formation
Trachoparia intermediaSp. novValidYuan et al.CambrianChanghia Formation
Trachoparia spinosaSp. novValidYuan et al.CambrianChanghia Formation
TriangulopygeGen. et comb. novValidHøyberget & BrutonCambrian

A member of the family Olenidae. The type species is "Olenus" humilis Phillips (1848); genus also includes "Sphaerophthalmus" majusculus Linnarsson (1880) and T. major (Lake, 1913).
Tsunyidiscus pengshuiensisSp novValidZhang & ClarksonCambrianAn eodiscoid trilobite, a species of Tsunyidiscus.
Ulania concavaSp. novValidYuan et al.CambrianChanghia Formation
Utagnostus primitivusSp. novValidYuan et al.CambrianChanghia FormationA member of the family Clavagnostidae.
Utagnostus sulcatusSp. novValidYuan et al.CambrianChanghia FormationA member of the family Clavagnostidae.
WoqishanaspisGen. et comb. novValidYuan et al.CambrianChanghia FormationA member of the family Ignotogregatidae. Genus includes W. trapezoidalis (Lin, 1984).
Wutaishanaspis yangjiazhuangensisSp. novValidYuan et al.CambrianChanghia Formation
Xenolorenzella granulosaSp. novValidYuan et al.CambrianChanghia Formation
YichengocephalusGen. et sp. novValidYuan et al.CambrianChanghia FormationA member of the family Solenopleuridae. Genus includes new species Y. latus.
Yongwolia quadrataSp. novValidYuan et al.CambrianChanghia Formation
Yushugouia ezhuangensisSp. novValidYuan et al.CambrianChanghia Formation
ZhangxiaspisGen. et sp. novValidYuan et al.CambrianChanghia FormationA member of the family Ptychopariidae. Genus includes new species Z. subcylindrica.
ZhongtiaoshanellaGen. et sp. novValidYuan et al.CambrianChanghia FormationA member of the family Crepicephalidae. Genus includes new species Z. brevispinosa.
Zhongweia convexaSp. novValidYuan et al.CambrianChanghia Formation
Zhujia yangjiazhuangensisSp. novValidYuan et al.CambrianChanghia FormationA member of Ptychopariida belonging to the family Anomocarellidae.
ZichuanaspisGen. et 2 sp. et comb. novValidYuan et al.CambrianChanghia FormationA member of the family Leiostegiidae. Genus includes Zichuanaspis acerius (Walcott, 1905) and Z. catinus (Resser & Endo in Endo & Resser, 1937), as well as new species Z. yangjiazhuangensis and possibly Z? latilimbata.

Others

NameNoveltyStatusAuthorsAgeUnitLocationNotesImages
Australimicola[128] Gen. et sp.ValidPaterson, García-Bellido and EdgecombeEarly CambrianThe Emu Bay Shale Konservat-LagerstätteAn artiopodan arthropod. The type species is Australimicola spriggi.
Kangacaris shui[129] Sp. novValidZhang, Fu & DaiLower CambrianChengjiang lagerstätteA species of Kangacaris.
Luohuilinella[130] Gen. et sp. novValidZhang, Fu & DaiLower CambrianChengjiang LagerstätteA possible member of Xandarellida. The type species is Luohuilinella rarus.
Nereocaris[131] Gen. et sp. novValidLegg et al.CambrianBurgess Shale FormationA bivalved arthropod, one of the basalmost known arthropods. The type species is Nereocaris exilis.
Pleuralata[132] Gen. et sp. novValidMcCoy, Strother and BriggsSilurian (Llandovery)Tuscarora FormationAn arthropod of uncertain phylogenetic affinities, a probable tracemaker of Arthrophycus alleghaniensis trace fossils. The type species is Pleuralata spinosa.
Scolopocryptops simojovelensis[133] Sp. novValidEdgecombe et al.MioceneA scolopocryptopid scolopendromorph centipede, a species of Scolopocryptops.
Sinoleperditia hamadai[134] Sp. novValidTanaka, Ono and MaedaEarly DevonianA member of Leperditicopida (a group of arthropods of uncertain affinities, classified as ostracods or excluded from this group by different authors), a species of Sinoleperditia.
Sinoleperditia huyeni[135] Sp. novValidTanaka, Phong and KomatsuEarly DevonianA member of Leperditicopida, a species of Sinoleperditia.
Squamacula buckorumSpeciesValidPaterson, García-Bellido and EdgecombeEarly CambrianThe Emu Bay Shale Konservat-LagerstätteAn artiopodan arthropod, a species of Squamacula.

Notes and References

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  3. Book: Jörg Wunderlich . 2012 . New fossil spiders (Araneae) of eight families in Eocene Baltic amber, and revisions of selected taxa . Jörg Wunderlich . Beiträge zur Araneologie, 7: Fifteen papers on extant and fossil spiders (Araneae) . 94–149 . http://joergwunderlich.de/Downloads/Beitr._Araneol._Band_7_(2012).pdf .
  4. George Poinar Jr. . Ron Buckley . 2012 . Predatory behaviour of the social orb-weaver spider, Geratonephila burmanica n. gen., n. sp. (Araneae: Nephilidae) with its wasp prey, Cascoscelio incassus n. gen., n. sp. (Hymenoptera: Platygastridae) in Early Cretaceous Burmese amber . Historical Biology . 24 . 5 . 519–525 . 10.1080/08912963.2011.640399. 2012HBio...24..519P . 86838688 .
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  6. Book: Jörg Wunderlich . 2017 . New and rare fossil spiders (Araneae) in mid Cretaceous amber from Myanmar (Burma), including the description of new extinct families of the suborders Mesothelae and Opisthothelae as well as notes on the taxonomy, the evolution and the biogeography of the Mesothelae . Jörg Wunderlich . Beiträge zur Araneologie, 10 . 72–279 . http://joergwunderlich.de/Downloads/Beitr._Araneol._Band_10_(2017).pdf .
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